Zarcosia cephalica ( Werner, 1962 ) Gompel, 2020

Gompel, Nicolas, 2020, Synopsis of the Asian species in the genus Zarcosia Collado & Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species, Zootaxa 4838 (2), pp. 151-178 : 156

Zarcosia cephalica ( Werner, 1962) , comb. nov.

( Figures 1 ; 2f ; 4d ; 5u )

Euglenes cephalicus Werner, 1962: 126 ; Holotype: Bishop Museum (Hawaii, USA); 1 paratype at the BMNH

Zarcosia cephalicus [sic!] ( Werner, 1962): Grzymala, 2016

Type locality. Malaysia: Selangor Darul Ehsan, Gombak District, Batu caves .

Distribution. Malaysia: Batu caves; Sarawak, Gunung Mulu National Park ( Figure 1 ).

Material examined. Malaysia: Sarawak, 4th Division, Gunung Mulu National Park, Deer Cave , (360 m) 04°07’ N 114°55’ E (P. M. Hammond & J. E. Marshall leg.) 6.V.1978 (1 ♂, 2 ♀, BMNH), 20.V.1978 (2 ♂, 1 ♀, BMNH; 1 ♂, 1 ♀, NGPC) ; Sarawak, 4th Division, Gunung Mulu National Park, Clearwater Cave , (360 m) 04°07’ N 114°55’ E (P. M. Hammond & J. E. Marshall leg.) 20.V.1978 (1 ♂, BMNH) . Republic of Indonesia: Aceh Selatan, Sawang, Tring Meuduro Tunong , Gua Kiciarean , Cave , (30 m) 03°23’35” N 97°07’26” E (L. Deharveng & A. Bedos leg.) 12.VII.1998 (1 ♂, 1 ♀, MNHN; 1 ♀, MZB), (J. Gibert leg.) 12.VII.1998 (1 ♀, MZB) ; Jawa Tengah, Cilacap, Nusakambangan Island, Goa Ratu , (70 m) cave, 07°43’ S 108°57’ (L. Deharveng & A. Bedos leg.) 17.II.2011 (1♀, NGPC) .

Figures

FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line.

FIGURE 2. continued next page.

FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm.

FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm.