Diakonoffiana laosensis Phewphanh and Pinkaew, 2017

Pinkaew, Nantasak & Phewphanh, Phonepadith, 2017, First report on the Olethreutini (Lepidoptera: Tortricidae: Olethreutinae) of Lao PDR with descriptions of two new species, Zootaxa 4358 (1), pp. 125-141 : 127-128

Diakonoffiana laosensis Phewphanh and Pinkaew , n.sp.

( Figs. 7–8 , 35, 37, 39–43)

Diagnosis. Diakonoffiana laosensis is superficially most similar to D. vindemians (Meyrick, 1912) in forewing shape and pattern, but differs in having a silvery rhomboidal mark on the costa. The male genitalia are also most similar to those of D. vindemians with a curved and clavate uncus and an elongate ovate valva with a dilated cucullus, but they can be distinguished from those of the latter by the sacculus bearing asymmetrical processes dorsally, each with a cluster of spiniform setae, and only the right valva bearing a strong, short thorn-like process. The female genitalia also are similar to those of D. vindemians in having an aciculate, complex sterigma with two lateral aciculate knobs, a long ductus bursae, and a pear-shaped corpus bursae with plate-shaped signa. The female genitalia of D. laosensis differ by having a very short ductus bursae and a larger corpus bursae.

Description. Head: Upper frons brown to dark brown, lower frons white mixed with brown; vertex light brown mixed with brown; labial palpus porrect (Fig. 35), first segment greyish brown, second segment slightly expanded apically, pale brown mixed with brown, third segment rather short, pale brown. Antenna reaching middle of forewing, brown. Thorax: Pronotal collar, tegulae, and mesonotum brown to dark brown, posterior crest brown; hindtibia with tuft of long, pale brown scale pencils on dorsal inner surface (Fig. 37). Forewing subrectangular, slightly broadened towards termen, length 6.1–7.0 mm in male (n = 11) ( Fig. 7 ), 7.0–7.3 mm in female (n = 4) ( Fig. 8 ); costal margin distinctly curved near base then slightly curved to apex, with two distinct silvery striae and a silvery rhomboidal mark medially in male, mark squarish in female; termen rather straight, with two small white spots, one below apex between R 4 and R 5 and second between R 5 and M 1, with a narrow, orange brown line between R 5 and M 3; tornus broadly rounded; forewing ground color greyish brown, with large, irregular, dark brown patch ventrobasally, with widened and curved band surrounding silvery rhomboidal mark, then extending along costa to termen, distal part with pinkish brown subcircular patch extending from R 5 to CuA 1. Underside pale brown. Hindwing brown, slightly paler at base, underside pale brown. Abdomen: Male genitalia (Fig. 39–41) with tegumen triangular, with prominent, sclerotized shoulders, apex with group of moderately dense sockets (setae lost during dissection); uncus moderately long, clavate, apex with a brush of dense setae, sparsely setose basally; socii membranous with a vertical row of setae; gnathos with lateral arms sclerotized, medially fused with prominent triangular subscaphium; vinculum a moderately wide band; juxta large, subtriangular; caulis short, and anellus forming membranous ring surrounding basal one-third of phallus, phallus rather small, subtriangular, wide at base and tapering to apex, without cornuti; valva club-shaped; saccullus with moderately dense setae basally, ventral margin with a group of dense setae, with two asymmetrical spiniform setae developed into a processes on basal edge of excavation, only right valva bearing strong, stout, rather short thorn-like process; cucullus dilated with brush of spiniform setae. Female genitalia (Fig. 42–43) with tergum VIII smooth except for lateral subtriangular extension with group of scale sockets; sternum VII with deeply U-shaped posterior margin, papillae anales bearing dense setae; ostium associated with complex, spinulose lamella postvaginalis (Fig. 42), with a vertically folded median structure above, and two small dome-shaped lateral knobs; colliculum sclerotized, ductus bursae very short, ductus seminalis arising at intersection with corpus bursae; corpus bursae pear-shaped, with two unequal, strongly curved, plate-shaped signa.

Holotype. ♂, LAOS, Vientiane Prov., Ban Vung Heuo I, Phou Khao Khouay N.P., 18 ̊21′12″N, 102 ̊48′85″E, ca. 740 m, 11 Jun 2016, P. Phewphanh, pp00193 (genitalia slide PP00093).

Paratypes. 10♂, 4♀, LAOS : Vientiane Prov., Ban Vung Heuo I, Phou Khao Khouay N.P., 18 ̊21′12″N, 102 ̊48′85″E, ca. 740 m, 28 Jan 2017, P. Phewphanh, pp00476 (♂ genitalia slide PP00183) . THAILAND: Prachinburi Prov., Khao Yai N.P., 14 ̊11′02″N, 101 ̊36′19″E, ca. 94 m, 18 June 2010, N. Pinkaew, np4477 (♂) , Khao Yai N.P., 14 ̊10′56″N, 101 ̊28′39″E, ca. 201 m, 13 May 2010, N. Pinkaew, np4082 (♀ genitalia slide NP1459), Khao Yai N.P., 14 ̊23′56″N, 101 ̊22′16″E, ca. 780 m, 9–12 Sep 2010, N. Pinkaew, np6722 (♂), N. Pinkaew, np6723 (♂), N. Pinkaew, np6724 (♂), Khao Yai N.P., 14 ̊21′49″N, 101 ̊24′37″E, ca. 1,140 m, 8 May 2010, N. Pinkaew, np3584 (♂), Khao Yai N.P., 14 ̊11′02″N, 101 ̊36′19″E, ca. 94 m, 24 Jan 2010, N. Pinkaew, np3403 (♂), Khao Yai N.P., 14 ̊11′02″N, 101 ̊36′19″ E, ca. 94 m, 15 Feb 2010, N. Pinkaew, np3464 (♂). Chanthaburi Prov., Khao Khitchakut N.P., 12 ̊50′21″N, 102 ̊07′15″E, ca. 73 m, 21 Feb 2012, N. Pinkaew, np7715 (♀ genitalia slide NP3096), Khao Khitchakut N.P., 12 ̊48′37″N, 102 ̊09′11″E, ca. 92 m, 6 June 2013, N. Pinkaew, np5842 (♂ genitalia slide NP1982) , Chanthaburi Prov., Ang-et Com. For. , 12 ̊36′04″N, 102 ̊19′50″E, ca. 33 m, 22–23 Dec 2011, N. Pinkaew, np8743 (♀ genitalia slide NP3095) , Trat Prov., Trat Agroforestry R. St. , 12 ̊23′43″N, 102 ̊40′32″E, ca. 30 m, 18–19 Aug 2012, N. Pinkaew, np5307 (♂ genitalia slide NP1563) . Nakhon Si Thammarat Prov., Khao Nan N.P., 08 ̊51′47″N, 99 ̊73′36″E, ca. 207 m, 19 Jan 2007, N. Pinkaew, np1973 (♀ genitalia slide NP1083) .

Distribution. Laos and Thailand.

Etymology. The specific epithet laosensis refers to Laos, the country of the holotype.

Figures

FIGURES 7–16. Wing patterns. 7. Diakonoffiana laosensis, n.sp. (holotype male) 8. Do. n.sp. (paratype female) 9. Dicephalarcha herbosa (Meyrick, 1909) 10. Dudua aprobola (Meyrick, 1886) 11. Dudua charadraea (Meyrick, 1909) 12. Dudua tetanota Meyrick, 1909 13. Gatesclarkeana idia Diakonoff, 1973 14. Hedya iophaea Meyrick, 1912 15. Lobesia acicula, n.sp. (holotype male) 16. Lobesia aeolopa Meyrick, 1907.