Nomada quadrimaculata (Robertson 1903)

Gibbs, Jason, Ascher, John S., Rightmyer, Molly G. & Isaacs, Rufus, 2017, The bees of Michigan (Hymenoptera: Apoidea: Anthophila), with notes on distribution, taxonomy, pollination, and natural history, Zootaxa 4352 (1), pp. 1-160 : 62

Nomada quadrimaculata (Robertson 1903) (ruficornis group: bidentate mandible) (New status, new state record)

Gnathias ovatus var. quadrimaculatus Robertson 1903: 175 [female lectotype, INHS].

County records: Alpena, Antrim, Clare, Gladwin, Oakland, Van Buren.

Notes. This species was originally described as Gnathias ovatus var. quadrimaculatus , but has not been in common usage, having been typically synonymized with the nominal species. JG makes this tentative determination based on examination of the lectotype. It differs from N. ovata in part by the more numerous five or six specialized setae on the apex of the hind tibia, but it is possible this character varies and is not reliable for species delimitation. Other putative forms of N. ovata , described by Robertson and listed above, differ in the number of these setae, in addition to their varying color patterns.

Material examined. Alpena Co.: (no locality), 2 Jun. 1951, R.R. Dreisbach (1 ♀ MSUC): Antrim Co.: Glacial Hills, Vandermark entrance, 44.987 -85.2417, 12 Jun. 2016, Rubus, K. Odanata, J. Brokaw, S. Naranjo (1 ♀ MSUC); Glacial Hills, Vandermark entrance, 44.987 -85.2417, 12 Jun. 2016, J. Gibbs (1 ♀ JBWM); Glacial Hills, Eckhardt entrance, 44.9906 -85.292, 12 Jun. 2016, Rubus, K. Odanata, J. Brokaw, S. Naranjo (2 ♀ JBWM); Clare Co.: (no locality), 20 Jun. 1950, R.R. Dreisbach (1 ♀ MSUC); Clinton Co.: Bath, 19 May 1956, R.L. Fischer (1 ♀ MSUC): Gladwin Co.: (no locality), 10–16 Jun. 1951, R.R. Dreisbach (1 ♀ MSUC); Mackinac Co.: (no locality), 7 Jun. 1960, R. & K. Dreisbach (1 ♀ MSUC): Oakland Co.: (no locality), 12 Aug. 1923 [illegible] (1 ♀ MSUC); Van Buren Co.: 3 mi. N Friendsville, 21 May 2005, J.K. Tuell et al. (1 ♀ MSUC: RI).