Eusarcus gemignanii (Mello-Leitão) Hara & Pinto-Da-Rocha, 2010

Hara, Marcos Ryotaro & Pinto-Da-Rocha, Ricardo, 2010, Systematic review and cladistic analysis of the genus Eusarcus Perty 1833 (Arachnida, Opiliones, Gonyleptidae) 2698, Zootaxa 2698, pp. 1-136 : 60-63

Eusarcus gemignanii (Mello-Leitão) comb. n., stat.n.

( Figs. 16, 46C,D, 51)

Pygophalangodus gemignanii Mello-Leitão 1931b: 83 (cit), 85, fig. 4; Ringuelet 1954: 1 (syst); 1955a: 294 (syst); 1959: 381 (rdesc), fig. 56; 1963: 49 (key); Galiano & Maury 1979: 322 (cat); Acosta et al. 1993: 29 (cit); Acosta 1993: 14 (cit). (ma holotype, 1 ma & 1 fe paratypes, “ Entre Ríos; Federal; C. Daguerre leg.; ii.1931; MACN 4755”; examined).

Pygophalangodus gemignanii gemignanii: Ringuelet 1955a: 294 (diag); 1957a: 20 (cat); 1959: 197 (cat), 380 (key), lam. XVII, figs. 2, 4; 1963: 49 (key); Acosta 1996a: 77 (cit); Maury & Pilati 1996: 2 (cit).

Pygophalangodus geminganii gemignanii (lapsus): Ringuelet 1959: 380 (rdesc).

Material examined: ARGENTINA. Entre Ríos: Federal, C. Daguerre leg., ii.1931, ma holotype , 1 ma & 1 fe paratypes ( MACN 4755 ) .

Diagnosis: E. gemignanii resembles E. caparaoensis + and E. manero +, which have a conical, short, blunt prolateral basal apophysis on the male trochanter IV, and can be distinguished by: The presence of PAM; ocularium next to anterior scutal margin; high, with two large tubercles; median spine of scutal area III absent; femur IV with a retrolateral row of enlarged, blunt tubercles; ventrally with a prolateral row of tubercles increasing in size distally, becoming high, pointed, 1 PVS, 1 large retrolateral apical tubercle.

Redescription: Male (holotype): Dorsum ( Fig. 16A,B): Measurements: SL 3.90; SMW 3.20; femur I 1.70; II 2.75; III 2.10; IV 2.80. Median paracheliceral projection smaller than PAM. Ocularium next to anterior scutal margin; high, with 2 large tubercles, 26 small ones. Carapace with 77 scattered tubercles. Scutal area I with 19–20 scattered tubercles on each side; II with 39; III with 40 and the median spine absent; IV with 44 tubercles. Posterior margin of dorsal scutum and free tergites I–III with a row of 21, 18, 17, 18 tubercles, respectively. Anal operculum with 36 tubercles.

Venter: Coxa I with 24 tubercles, median and distal ones largest; II with 33–35, distal ones largest; III–IV irregularly tuberculate.

Chelicera: Segment I with 1–2 tubercles.

Pedipalpus: Trochanter dorsally inflated, with 2 tubercles. Femur dorsally with 5 aligned tubercles; ventrally with 1 large pointed basal tubercle, 5 small aligned ones. Tibial setation: Prolateral IiIi/IiI, retrolateral IiIi. Tarsal setation: Prolateral IIi, retrolateral IiIi/iIi.

Legs ( Fig. 16C–J): Coxa II with 2 tubercles on retrolateral margin; IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, long, blunt, curved posteriad and ventrad, swollen ventro-apically. Trochanters I–IV with scattered tubercles; IV prolaterally with 1 basal apophysis, this conical, short, blunt, swollen in the middle. Femur I with small PDS, reduced RDS; ventrally with 2 rows of slightly enlarged tubercles; II with small PDS, RDS; III sinuous, with small PDS, RDS; ventrally with 2 rows of tubercles increasing in size distally, prolateral ones enlarged; 1 large prolateral apical tubercle; IV sinuous, with medium PDS, small RDS; with a retrolateral row of enlarged, blunt tubercles; ventrally with a prolateral row of tubercles increasing in size distally, becoming high, pointed, 1 PVS, 1 large retrolateral apical tubercle. Tibia III ventrally with 2 rows of tubercles increasing in size subapically, becoming high, pointed; IV ventrally with 2 rows of tubercles increasing in size distally, prolateral ones enlarged, 2 apical spines. Tarsal segmentation: 5, 6, 6, 6.

Penis (MACN 4755; Fig. 46C,D): Stylus smooth, apex slightly inflated and with angular medio-ventral projection. Ventral process of glans curved; apex blunt. Ventral plate with concave sides; with 3 pairs of curved distal setae; 1 pair of short, straight median setae; 4 pairs of straight basal setae.

Coloration: Brown; dark brown on prolateral apical apophysis of coxa IV and from distal part of trochanter IV to basal half of metatarsus IV, distal half of metatarsus IV light brown. Light brown on distal portions of all podomeres.

Female (paratype; MACN 4755): Dorsum: Measurements: SL 3.75; SMW 3.10; femur I 1.60; II 2.60; III 1.95; IV 2.50. Median paracheliceral projection moderately enlargedr than PAM. Posterior margin of dorsal scutum with 10 scattered tubercles and a row of 24 tubercles. Pedipalpus: Tibial setation: Prolateral and retrolateral IiIi. Tarsal setation: Prolateral IIi, retrolateral IiIi. Legs: Coxa IV densely tuberculate, with 1 oblique prolateral apical apophysis, this conical, short (shorter than in male), pointed. Trochanter I with a large ventro-basal tubercle; IV with 1 large retrolateral apical tubercle. Femur I ventrally with 2 rows of similar-sized tubercles; II with small PDS, reduced RDS; III; ventrally with a prolateral row of tubercles increasing in size distally, 1 large prolateral apical tubercle; IV with a retrolateral row of similar-sized tubercles; ventrally with a prolateral row of tubercles increasing in size distally, 1 small, blunt PVS. Tibia III with 2 ventral rows of tubercles slightly increasing in size subapically; IV with 2 ventral rows of similar-sized tubercles, without apical spines. Tarsal segmentation: 5, 6, 6, 6. Coloration: Brown; light brown on apex of metatarsi and tarsi.

Variation in males (n=2): Dorsum: Measurements: SL 3.60–3.90; SMW 2.95–3.20; femur I 1.50–1.70; II 2.45–2.75; III 1.90–2.10; IV 2.60–2.80. Median paracheliceral projection smaller to larger than PAM. Pedipalpus: Tibial setation: Prolateral IiIi/IiI. Tarsal setation: Retrolateral IiIi/iIi. Trochanter I with or without 1 large ventro-basal tubercle; IV with or without 1 large retrolateral apical tubercle; femur I with reduced or small PDS; II with reduced or small PDS, RDS; tibia IV with 2 ventral rows of tubercles increasing in size distally, prolateral ones slightly enlarged.

Type locality: Argentina, Entre Ríos, Federal .

Geographical distribution ( Fig. 51): Known only from the type locality.