Perrona styriaca ( Hilber, 1879 ), 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 131-132

publication ID	https://doi.org/10.11646/zootaxa.5123.1.1
publication LSID	lsid:zoobank.org:pub:036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A
persistent identifier	https://treatment.plazi.org/id/039487D1-FF00-FF2B-FFBA-FB006930F9E7
treatment provided by	Plazi (2022-03-30 20:52:14, last updated by Valdenar 2024-02-29 15:27:11)
scientific name	Perrona styriaca ( Hilber, 1879 )
status	comb. nov.

Treatment

Perrona styriaca ( Hilber, 1879) View in CoL nov. comb.

Figs 40A View FIGURE 40 1 –A View FIGURE 1 3 View FIGURE 3 , B 1 –B View FIGURE 1 3 View FIGURE 3 , 4V View FIGURE 4 , 5 View FIGURE 5 , 7 View FIGURE 7

* Pleurotoma (Clavatula) styriaca Auing. in coll.— Hilber 1879: 19, pl. 3, figs 6–7.

Pleurotoma (Clavatula) styriaca Auing. — Hoernes & Auinger 1891: 348 (partim), pl. 47, figs 4–5 [non figs 6–9 = Clavatula pilleri nov. sp.; non fig. 10 = Perrona lydiae (Hoernes & Auinger, 1879) View in CoL ].

Clavatula styriaca Hilber View in CoL , sp. 1879— Glibert 1960: 42.

Type material. Lectotype: NHMW 1846/0037/0309a, Pöls ( Austria), SL: 21.8 mm, MD: 9.3 mm, illustrated in Hilber (1879, pl. 3, figs 7a–c) and Hoernes & Auinger (1891, pl. 47, figs 5a–c), designated by Švagrovský (1982: 412), figs 40A 1 –A 3, 4 V. Paralectotypes: NHMW 1846/0037/0309b, Pöls ( Austria), SL: 40.5 mm, MD: 16.1 mm, illustrated in Hilber (1879, pl. 3, fig. 6) and Hoernes & Auinger (1891, pl. 47, fig. 4, figs 40B 1 –B 3; NHMW 1949/0005/0051f, Pöls ( Austria), SL: 44.7 mm, MD: 18.8 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 8); NHMW 1949/0005/0051g, Pöls ( Austria), SL: 49.9 mm, MD: 18.2 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 9); NHMW 1861/0001/0266, 2 spec., Pöls ( Austria).

Revised description. Shell small, solid, moderately broad fusiform with high spire; apical angle 30–33°. Protoconch not preserved. Teleoconch of ten whorls. Early teleoconch whorls flat-sided with two smooth, narrow adsutural spiral cords separated by broad mid-portion bearing fine, opisthocline axial riblets. Riblets fading out on fourth or fifth teleoconch whorls. Later whorls concave, with periphery at abapical suture. Prominent subsutural collar bearing widely-spaced pointed tubercles placed at mid-collar, developed from eighth whorl. Suprasutural cord almost completely obscured by succeeding whorl. Spiral sculpture of delicate spiral threads weakening on last whorl. Suture narrowly incised, superficial, linear. Last whorl ~60% of total height; of medium width. Weakly coronate subsutural collar bearing 10–12 pointed tubercles. Subsutural ramp wide, concave, delimited by angular shoulder placed at level of insertion of outer lip. Base placed high, in short distance below shoulder, weakly angled, strongly constricted. Weak siphonal fasciole. Entire surface covered in fine spiral cords of alternating strength and slightly stronger over base and fasciole. Aperture wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide and deep, asymmetrically U-shaped, with apex mid-ramp; siphonal canal moderately long, narrow, slightly deflected to the left, shallowly notched at tip. Columella strongly excavated, twisted at fasciole, smooth. Columellar and parietal callus moderately thickened, adherent, forming broad callus rim.

Discussion. This species was until now placed in the genus Clavatula by all authors. However, the commashaped axial riblets on early teleoconch whorls suggest a placement in Perrona . Perrona styriaca ( Hilber, 1879) is surprisingly rare and the wide distribution suggested by the literature records is based on numerous misidentifications. The concept of this species was also blurred by Hoernes & Auinger (1891) by including an aberrant specimen of Perrona lydiae ( Hoernes & Auinger, 1891) and by including specimens of Megaclavatula pilleri nov. sp. from Baden-Sooss ( Austria) in their description. Perrona lydiae , however, is clearly separated from P. styriaca by its swollen adsutural cords, higher spire and more slender outline. Megaclavatula pilleri is distinguished by its tripartite sculpture on the early teleoconch whorls, as opposed to the comma-shaped axial riblets typical of Perrona species. Several authors placed small sized and subadult shells of Megaclavatula laevigata ( Eichwald, 1830) in P. styriaca (e.g., Švagrovský 1958, Strausz 1966), which all differ from P. styriaca in having tripartite early teleoconch whorl sculpture. Similarly, the single specimen from Korytnica, reported by Bałuk (2003) as P. styriaca has a reduced peribasal spiral cord and differs from P. styriaca also in its shorter spire and wider apical angle. Specimens from the Carpathian Foredeep, referred to as Clavatula styriaca by Friedberg (1912, 1938) and Bałuk (1970) were described as Clavatula kowalewskii by Bałuk (2003) [= Megaclavatula antoniae ( Hoernes & Auinger, 1891) ], which differs from P. styriaca in its slender fusiform outline, a conical spire and strongly reduced sculpture on the subsutural collar.

Paleoenvironment. Shallow marine, inner neritic, based on the mollusc assemblage from Pöls ( Austria) (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Styrian Basin: Pöls ( Austria) ( Hoernes & Auinger 1891).

References

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Baluk, W. (2003) Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part IV - Turridae. Acta Geologica Polonica, 53, 29 - 78. [https: // geojournals. pgi. gov. pl / agp / article / view / 9720 / 8257]

Eichwald, E. (1830) Naturhistorische Skizze von Lithauen, Volhynien und Podolien in Geognostisch-Mineralogischer, Botanischer und Zoologischer Hinsicht. Wilna, Voss, 255 pp. [https: // reader. digitale-sammlungen. de / de / fs 1 / object / display / bsb 10691419 _ 00005. html]

Friedberg, W. (1912, 1928) Mieczaki Miocenskie ziem Polskich. (Mollusca Miocaenica Poloniae). Czesc I. Slimaki i Lodkonogi (Pars I. Gastropoda et Scaphopoda). Muzeum Imienia Dzieduszyckich We Lwowie. Vols. 1 & 5. Polskie Towarzystwo Geologiczne, Krakoiw, pp. 113 - 239, pls. 6 - 14 (1912) & pp. 441 - 631, pls. 27 - 38 (1928). [https: // rcin. org. pl / miiz / dlibra / publication / 4795 / edition / 29891 / content]

Friedberg, W. (1938) Katalog mego zbioru mieczakow miocenskich Polski (Katalog meiner Sammlung der Miozanmollusken Polens). Memoires de l'Academie polonaise des sciences et des lettres, Classe des sciences mathematiques et naturelles. Serie B, Sciences naturelles, 12, 1 - 164.

Glibert, M. (1960) Les Conacea Fossiles du Cenozoique Etranger des collections de l'Institut Royal des Sciences Naturelles de Belgique. Memoires de l'Institut Royal des Sciences Naturelles de Belgique, 64, 1 - 133. [https: // core. ac. uk / download / pdf / 35118094. pdf]

Hilber, V. (1879) Neue Conchylien aus den mittelsteierischen Mediterranschichten. Sitzungsberichte der kaiserlichen Akademie der Wissenschaften, der Mathematisch-Naturwissenschaftlichen Classe, 79, 416 - 464. [https: // www. zobodat. at / pdf / SBAWW _ 79 _ 0416 - 0464. pdf]

Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]

Kokay, J. (1966) Geologische und palaontologische Untersuchung des Braunkohlengebietes von Herend-Marko (Bakony- Gebirge, Ungarn). Geologica Hungarica, Series Palaeontologica, 36, 1 - 149. [https: // epa. oszk. hu / 02900 / 02989 / 00029 / pdf /]

Mikuz, V. (1998) Turridae (Neogastropoda) iz srednjemiocenskih badenijskih plasti Slovenije. (Turrids (Neogastropoda) from Middle Miocene Badenian beds of Slovenia). Geologija, Ljubljana, 40, 65 - 101. [1997, http: // www. geologija-revija. si / dokument. aspx? id = 795] https: // doi. org / 10.5474 / geologija. 1997.002

Mikuz, V. (2009) Miocene gastropods from the vicinity of Sentjernej and from other localities in the Krka Basin, Slovenia. Folia Biologica et Geologica, 50, 5 - 69. [http: // www. sazu. si / uploads / files / 57 dfbe 71 e 126 b 1 a 75 cebe 90 f / 420 - 122 - 0. pdf]

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Strausz, L. (1954) Varpalotai felso-mediteean csigak. Geologica Hungarica, 25, 1 - 150. [http: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954 _ 085 - 125. pdf]

Strausz, L. (1966) Die Miozan-Mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp.

Svagrovsky, J. (1958) Miocenne Pleurotomidae zapadokarpatskych panvi (Miozane Pleurotomidae der westkarpatischen Becken). Acta Geologica et Geographica Universitatis Comenianae, Geologica, 1, 5 - 56.

Svagrovsky, J. (1982) Gastropoda, Prosobranchia. Teil 2. Neogastropoda des oberen Badeniens von Borsky Mikulas (NO-Teil des Wiener Beckens) und ihre stratigraphische Bedeutung. Geologicky Zbornik; Geologica Carpathica, 33 / 4, 383 - 435. [http: // www. geologicacarpathica. com / browse-journal / archive- 1950 - 1996 / #]

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Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 40 A1–A3. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309a, Pöls (Austria). B1–B3. Perrona styriaca (Hilber, 1879), paralectotype, NHMW 1846/0037/0309b, Pöls (Austria). C1–C3. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). D1–D3, Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). E1–E3. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0107b, Buituri (Romania). F1–F3. Perrona ursulae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0107a, Buituri (Romania).