Perrona wanzenboecki, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 133-135

Perrona wanzenboecki nov. nom.

Figs 40C 1 –C 3 , 4Y , 5 , 7

* Pleurotoma concinna n.f.— Handmann 1883: 171 [non Pleurotoma concinna Scacchi, 1836 ].

Perronea [sic] (Perronea) [sic] concinna Handmann — Mariani & Papp 1966: 144, pl. 1, figs 11–12 [non Pleurotoma concinna Scacchi, 1836 ].

Type material. Neotype: NHMW 2021 /0158/0001, SL: 23.2 mm, MD: 9.0 mm, St. Veit an der Triesting ( Austria), figs 40C 1 –C 3. The syntypes from St. Veit an der Triesting ( Austria) have been stored in the Kollegium Kalksburg in Vienna, a Catholic private school. One syntype has been illustrated in Mariani & Papp (1966, pl. 1, fig. 11) but this specimen is lost. Therefore , we designate a neotype from the type locality.

Type locality. St. Veit an der Triesting ( Austria), Vienna Basin .

Type stratum. Sandy nearshore equivalent of the Baden Formation.

Age. Middle Badenian (late Langhian).

Etymology. In honor of Gerhard Wanzenböck (Gainfarn, Austria), fossil collector and friend of the first author.

Revised description. Shell medium-sized, moderately broad fusiform with relatively short, conical spire. Protoconch largely destroyed, terminating with broad, strongly convex whorl. Teleoconch of seven whorls. First three teleoconch whorls faintly concave with narrow subsutural spiral cord and close-set, strongly opisthocline, comma-shaped axial riblets; no suprasutural cord. Sculpture changing on fourth teleoconch whorl; subsutural cord becomes moderately broad, rounded collar, whorl profile weakly concave below, bearing six narrow, flattened spiral cords. Suture narrowly incised, superficial, linear. Last whorl ~70% of total height, ovate. Subsutural collar broad, rounded, weakly swollen, sharply delimited. Subsutural ramp, shoulder and base not delimited. Profile below collar evenly rounded, strongly constricted at base. Siphonal fasciole poorly developed. Fine spiral cords over ramp, obsolete at mid-whorl, reappearing on abapical half of base, strengthening slightly towards and over fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately narrow and deep, asymmetrically U-shaped with apex mid-ramp. Siphonal canal long, straight, narrow. Columella weakly excavated, very slightly twisted at fasciole. Columellar and parietal callus thickened, well demarcated from base, forming broad rim.

Discussion. Handmann (1883) was unaware that his Pleurotoma concinna was variously preoccupied by Scacchi, (1836: 12), Adams (1852:140) and Dunker (1857: 356) (see Tucker 2004: 227). Therefore, we propose Clavatula wanzenboecki as replacement name. Perrona wanzenboecki nov. nom. is unique amongst its congeners in not having a suprasutural band developed and not having any of the major cords on the last whorl developed, so that the last whorl profile is evenly rounded. It is reminiscent of Tomellana dulaii nov. nom. and Megaclavatula nemethi ( Kovács & Vicián, 2021) in general shape, but is much smaller. In addition, T. dulaii has a slightly angulated subsutural collar and M. nemethi is slightly more slender and has tripartite early teleoconch whorl sculpture. Tomellana jouannetii ( Des Moulins, 1842) is also very similar, but is much larger, has a higher spire and lacks prominent spiral cords below the subsutural collar (see Peyrot 1931, pl. 8, figs 43, 61, 62, 66, 71). Moreover, both of Tomellana species compared above lack the comma-shaped axial riblets on the neanic whorls, typical of Perrona .

Paleoenvironment. Deposits and faunas from St. Veit an der Triesting represent an intertidal lagoon with mudflats and scattered oyster beds (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: St. Veit an der Triesting ( Austria) ( Mariani & Papp 1966).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 40 A1–A3. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309a, Pöls (Austria). B1–B3. Perrona styriaca (Hilber, 1879), paralectotype, NHMW 1846/0037/0309b, Pöls (Austria). C1–C3. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). D1–D3, Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). E1–E3. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0107b, Buituri (Romania). F1–F3. Perrona ursulae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0107a, Buituri (Romania).