Striopusionella szekelyhidiae ( Kovács & Vicián, 2021 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 142

Striopusionella szekelyhidiae ( Kovács & Vicián, 2021) nov. comb.

Figs 42B 1 –B 2 , 4 Ab, 5, 7

* Clavatula szekelyhidiae n. sp. — Kovács & Vicián 2021: 145, pl. 3, figs 7–10.

Type material. Holotype: HNHM, PAL 2021.19 .1, SL: 65.0 mm, MD: 25.6 mm, Letkés ( Hungary), illustrated in Kovács & Vicián (2021, pl. 3, figs 9–10); figs 42B 1 –B 3, 4 Ab . Paratype: NHMW 2021 /0002/0002, SL: 53.5 mm , MD: 19.4 mm, Letkés ( Hungary), illustrated in Kovács & Vicián (2021, pl. 3, figs 7–8) .

Revised description. Shell moderately large, solid, moderately slender fusiform with high conical spire; apical angle ~35°. Protoconch not preserved. Teleoconch of eleven whorls. Early teleoconch whorls with tripartite sculpture, weakly concave with two beaded adsutural cords, separated by wide concavity with weak central cord of delicate beads. Later teleoconch whorls broadly and weakly concave, subsutural collar narrow, weak, bearing small, wide-spaced tubercles, subobsolete in some specimens, suprasutural tubercles almost completely obscured by succeeding whorl. Entire shell covered by raised, narrow convex spiral cords, separated by interspaces of roughly equal width. Suture narrowly impressed, finely undulating. Last whorl 60% of total height. Subsutural collar very poorly delimited. Subsutural ramp broad, weakly concave, delimited and angled by narrow weakly tuberculose shoulder cord. Straight sided below, strengthened by very weakly tuberculose peribasal cord. Base moderately constricted, weakly concave in profile. Sinuous, poorly delimited axial folds starting on the abapical half of ramp and extending to base. Siphonal fasciole poorly delimited, strongly rounded and twisted, forming wide, deep pseudoumbilicus in fully adult specimens. Sculpture below shoulder cord of narrow irregularly beaded cords of roughly alternate strength. Aperture moderately wide, ovoid pyriform. Outer lip not thickened, smooth within. Anal sinus very wide, very shallow with apex mid-ramp. Siphonal canal short, narrow. Columella weakly excavated, smooth. Columellar and parietal callus strongly thickened, sharply delimited, moderately broad.

Discussion. We are not aware of comparable species in the Paratethyan fauna. ‘ Clavatula ’ occidentalis Peyrot, 1931, from the Langhian of Saubrigues ( France), displays a similar sculpture, but differs from the Paratethyan species in its shorter spire, longer siphonal canal and more concave spire whorls. Its anal canal is much deeper and therefore a closer relation with Striopusionella szekelyhidiae ( Kovács & Vicián, 2021) is unlikely (see Peyrot 1931, pl. 5, figs 10, 18). Similarly, ‘ Clavatula ’ saubrigniana ( Grateloup, 1832), from the Langhian of Saubrigues ( France), is similarly slender in outline, but has a deep anal sinus (see Peyrot, 1931, pl. 5, figs 21, 26, 27).

Paleoenvironment. The occurrence at Letkés ( Hungary) suggests a preference for inner neritic environments in the vicinity of corals ( Kovács & Vicián 2013).

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Letkés ( Hungary) ( Kovács & Vicián 2021).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 42 A1–A3. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). B1–B2. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). C1–C2. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). D1–D2. Tomellana praecursor (Schaffer, 1912), lectotype, KM Inv. No. F/1129, Maigen (Austria). E. Tomellana praecursor (Schaffer, 1912), paralectotype, KM Inv. No. F/1130, Maigen (Austria). F. ‘Clavatula’? czarnockii (Bałuk, 2003), holotype. Catalogue No. 28.II. 179, Korytnica (Poland). F1–F3. Tomellana aueri nov. sp., holotype, NHMW 1874/0033/0009a, Bad Vöslau (Austria). G1–G3. Tomellana aueri nov. sp., paratype, NHMW 1874/0033/0009b, Bad Vöslau (Austria). H. Tomellana aueri nov. sp., paratype, NHMW 2013/0078/0436, Baden (Austria).