Tomellana praecursor (Schaffer, 1912) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 147

Tomellana praecursor (Schaffer, 1912) nov. comb.

Figs 42D 1 –D 2 , E, 5

* Pleurotoma (Perrona) semimarginata Lam. var. praecursor Schff. — Schaffer 1912a: 135: pl. 49, figs 18–20.

Clavatula (Perrona) cf. consimilis (Bellardi) — Rutsch 1929; 14, pl. 1, fig. 1 [non ‘ Clavatula ’ consimils ( Bellardi, 1877)].

P [errona]. (P [errona]) semimarginata praecursor (Schff) — Sieber 1958a: 157.

Clavatula (Perrona) semimarginata ( Lamarck, 1822) — Hölzl 1958: 264, pl. 21, fig. 17 [non Tomellana semimarginata ( Lamarck, 1822) ].

Perrona semimarginata paulucciae Morlot — Steininger et al., 1971 b: 163 [nomen nudum].

Perrona (Perrona) semimarginata ( Lamarck, 1822) — Steininger et al. 1971: 405, pl. 11, fig. 4 [non Tomellana semimarginata ( Lamarck, 1822) ].

Perrona (Perrona) semimarginata praecursor (Schaffer) — Mandic & Steininger 2003: 274.

Clavatula semimarginata ( Lamarck, 1822) — Fözy & Leél-Össy 1985a: 121, pl. 3, fig. 6 [non Tomellana semimarginata ( Lamarck, 1822) ].

Clavatula intermedia miopercostata Sacco, 1904 — Fözy & Leél-Össy 1985a: 121 [non ‘ Clavatula ’ miopercostata ( Sacco, 1890)].

Clavatula semimarginata ( Lamarck, 1822) — Fözy & Leél-Össy 1985b: 184, pl. 3, fig. 7 [non Tomellana semimarginata ( Lamarck, 1822) ].

Clavatula intermedia miopercostata Sacco, 1904 — Fözy & Leél-Össy 1985b: 184 [non ‘ Clavatula ’ miopercostata ( Sacco, 1890)].

Clavatula semimarginata Lamarck — Ionesi & Nicorici 1994: 60, pl. 3, figs 17–18 [non Tomellana semimarginata ( Lamarck, 1822) ].

Type material. Lectotype designated herein: KM Inv. No. F /1129, SL: 47.1 mm, MD: 17.9 mm, Maigen ( Austria), illustrated in Schaffer (1912a, pl. 49, fig. 18), figs 42D 1 –D 2 . Paralectotypes: KM Inv. No. F/1130, SL: 26.4 mm , MD: 11.6 mm, Maigen ( Austria), illustrated in Schaffer (1912a, pl. 49, fig. 19), figs 42E ; KM Inv. No , F/1131 SL: 29.8 mm , MD: 12.5 mm, Maigen ( Austria), illustrated in Schaffer (1912a, pl. 49, fig. 20) .

Revised description. Shell moderately large, moderately broad fusiform with conical spire and high last whorl; apical angle ~40°. Protoconch and early teleoconch whorls not preserved. Early spire whorls weakly concave with weakly swollen subsutural collar and spiral row of broad, low, slightly opisthocline beads along suprasutural cord. Beads fading out on later teleoconch whorls. Entire shell covered by low, flat spiral cords separated by narrow interspaces. Suture indistinct, narrowly impressed. Last whorl 65% of total height, ovoid; subsutural collar rounded, weakly developed. Subsutural ramp of medium width, concave, poorly delimited by somewhat inflated, rounded shoulder marking periphery, convex below. Base weakly delimited, slightly angular, moderately constricted.Aperture narrowly pyriform. Outer lip incomplete. Anal sinus modertately wide and deep, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal long, narrow, straight with weakly notched tip.

Discussion. This species was established by Schaffer (1912a) as subspecies of Perrona semimarginata auctores [= Tomellana aueri nov. sp.], but differs from that species greatly in its shorter spire, wider apical angle, fewer teleoconch whorls, and the presence of a row of suprasutural beads. Tomellana praecursor is reminiscent of Tomellana dulaii nov. sp., from the Badenian of Nemeşeşti ( Romania) in general shape, but differs in its beaded suprasutural cord, weaker subsutural collar and shorter siphonal canal.

Paleoenvironment. The occurrence at Maigen ( Austria) suggests a preference for inner neritic environments with sandy bottoms (own data).

Distribution in Central Paratethys. Eggenburgian (early Miocene): North Alpine Foreland Basin: St. Gallen ( Switzerland) (Rutsch, 1926), Kaltenbachgraben ( Germany) ( Hölzl 1958); North Alpine-Carpathian Foreland Basin: Fels am Wagram, Maigen ( Austria) (Steiniger 1963), Eastern Mátra Mountains: Darnó-hill ( Hungary) ( Fözy & Leél-Össy 1985a, b); Dacian Basin: Crivienni-Pătârlagele ( Romania) ( Ionesi & Nicorici 1994).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 42 A1–A3. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). B1–B2. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). C1–C2. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). D1–D2. Tomellana praecursor (Schaffer, 1912), lectotype, KM Inv. No. F/1129, Maigen (Austria). E. Tomellana praecursor (Schaffer, 1912), paralectotype, KM Inv. No. F/1130, Maigen (Austria). F. ‘Clavatula’? czarnockii (Bałuk, 2003), holotype. Catalogue No. 28.II. 179, Korytnica (Poland). F1–F3. Tomellana aueri nov. sp., holotype, NHMW 1874/0033/0009a, Bad Vöslau (Austria). G1–G3. Tomellana aueri nov. sp., paratype, NHMW 1874/0033/0009b, Bad Vöslau (Austria). H. Tomellana aueri nov. sp., paratype, NHMW 2013/0078/0436, Baden (Austria).