Clavatula ’ atatuerki, Lamarck, 1801

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 26-27

‘ Clavatula ’ atatuerki nov. sp.

Figs 11E 1 –E 2 , F 1 –F 2 , 5

Clavatula olgae ( Hoernes & Auinger, 1891) — Landau et al. 2013 (partim): 286, pl. 48, figs 2–4, pl. 50, fig. 8 [non ‘ Clavatula ’ olgae ( Hoernes & Auinger 1891) ] [non Landau et al. 2013, pl. 48, figs 5–6 = ‘ Clavatula ’ ariejansseni nov. sp.] .

Type material. Holotype: NHMW 1847 /0058/1216, Seyithasan ( Turkey), SL: 41.4 mm, MD: 14.8 mm, illustrated in Landau et al. (2013, pl. 48, figs 3a–3b), figs 11E 1 –E 2 . Paratype: NHMW 1847 /0058/1217, Seyithasan ( Turkey), SL: 37.4 mm , MD: 13.5 mm, illustrated in Landau et al. (2013, pl. 48, figs 4a–4b), figs 11F 1 –F 2.

Type locality. Seyithasan ( Turkey), Karaman Basin .

Type stratum. Silt and clay of the Týrtar Formation.

Age. Middle Miocene, late Serravallian.

Etymology. Referring to Kemal Atatürk (1871–1938), first president of Turkey.

Diagnosis. Medium-sized, slender to moderately slender fusiform shell with elevated tuberculate adsutural cords separated by smooth concave mid-portion. Base with weak perifasciolar cords. Siphonal canal moderately long.

Description. Shell medium-sized, solid, slender to moderately slender fusiform; apical angle 37°. Protoconch not preserved. Early teleoconch spire whorls slightly concave, with bipartite sculpture of weak subsutural cord and beaded suprasutural cord, midsection deeply concave. On later whorls subsutural collar somewhat elevated, weakly delimited, bearing short, widely-spaced spines and one or two weak spiral threads. Mid-whorl portion smooth, broadening abapically, suprasutural cord bearing close-set, axially elongated tubercles. Last whorl 63– 66% total height, subsutural collar as described above, subsutural ramp broad, strongly concave, delimited by roundly angled shoulder bearing small, close-set, comma-shaped tubercles. Convex below, moderately constricted at base. Sculpture below shoulder of two to three primary cords of irregular strength, uppermost cord nodular, very irregular weak axial sculpture extends abapically from nodules beading spiral cords. Aperture small, ovate. Anal sinus moderately narrow, very deep, asymmetrically U-shaped. Weak inner lirae deep inside aperture. Columella moderately excavated in adapical third, weakly twisted. Siphonal canal moderately long, slightly twisted.

Discussion. This species was identified by Landau et al. (2013) as Clavatula olgae ( Hoernes & Auinger, 1891) , but ‘ Clavatula ’ olgae differs in the presence of spiral sculpture on the teleoconch whorls, the more prominent beads on the adsutural cords and the more prominent inner lirae. The Turkish species seems to be closer to the Paratethyan ‘ Clavatula ’ camillae ( Hoernes & Auinger, 1891) . Both species develop a marked and smooth mid-whorl concavity and comparatively small and reduced beads on the adsutural cords. Nevertheless, ‘ Clavatula ’ camillae is clearly distinguished from ‘ C. ’ atatuerki nov. sp. by its shorter siphonal canal and very prominent, tubercular perifasciolar cords.

‘ Clavatula ’ santhai Kovács & Vicián, 2021 differs in it much higher spire and the slowly contracting base. The specimens illustrated by Landau et al. (2013, pl. 48, figs 5–6) are excluded from ‘ Clavatula ’ atatuerki. Based on its tripartite early whorls it represents a species of the ‘ Clavatula ’ interrupta group and is described herein as ‘ Clavatula ’ ariejansseni nov. sp.

Paleoenvironment. Inner neritic environments ( Landau et al. 2013).

Proto-Mediterranean Sea: Serravallian (middle Miocene): Karaman Basin: Lale,Akboðazi, Akpýnar-Pýnarlar Yaylasý, Seyithasan, Road to Mut, 27 km south of Karaman ( Turkey) ( Landau et al. 2013).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 11 A1–A3. Clavatula sorini nov. sp., holotype, NHMW 1865/0001/0224, Lăpugiu de Sus (Romania). B1–B3. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania).C1–C3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania).D1–D3. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0042b, Lăpugiu de Sus (Romania), E1–E2. ‘Clavatula’ atatuerki nov. sp., holotype,. NHMW 1847/0058/1216, Seyithasan (Turkey). F1–F2. ‘Clavatula’ atatuerki nov. sp., paratype, NHMW 1847/0058/1217, Seyithasan (Turkey). G1–G2. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).