Clavatula ’ jarzynkae

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 42-43

‘ Clavatula ’ jarzynkae nov. nom.

Figs 15D 1 –D 3 , E 1 –E 3

Clavatula Clarae —Hoernes 1891: 129 [nomen nudum].

Pleurotoma (Clavatula) Clarae nov. form.— Hoernes & Auinger 1891: 342, pl. 45, figs 9a–c [non Pleurotoma clarae TenisonWoods, 1880].

Clavatula laevigata Eichw. var., aff. Cl. Clarae R. H. I A.—Friedberg 1928: 569, text-fig. 81 left.

Clavatula auingeri n. n.— Finlay 1927: 515 [nov. nom. pro Pleurotoma (Clavatula) clarae Hoernes & Auinger, 1891 , non Pleurotoma clarae Tenison-Woods, 1880 ] [non Hilber, 1879].

Type material. Lectotype designated herein: NHMW 1949/0005/0043, Grund ( Austria), SL: 47.2 mm, MD: 22.9 mm, syntype of Pleurotoma (Clavatula) clarae Hoernes & Auinger, 1891 , illustrated in Hoernes & Auinger (1891, pl. 45, figs 9a–c), figs 15D 1 –D 3. Paralectotypes: NHMW 1855/0002/0021, SL: 48.9 mm, MD: 21.8 mm, Grund ( Austria), figs 15E 1 –E 3; NHMW 1855/0002/0021, 2 spec., Grund ( Austria); NHMW 1855/0002/0021, 2 spec., Grund ( Austria).

Type locality. Grund ( Austria), North Alpine-Carpathian Foreland Basin .

Type stratum. Silty sand of the Grund Formation.

Age. Middle Miocene, early Badenian (Langhian).

Etymology. In honor of Agata Jarzynka (Polish Academy of Sciences, Kraków, Poland) in appreciation for her kind help with the search for type specimens from Poland and Ukraine.

Diagnosis. Large, solid, Clavatulidae with broad conical spire, prominent nodes on subsutural cord and long siphonal canal, differing from similar species by the absence of spines.

Description. Shell moderately large, solid, with broad conical spire; apical angle ~50°. Protoconch and early teleoconch not preserved. Teleoconch of more than eight whorls. Early teleoconch whorls with very narrow, weakly beaded subsutural and suprasutural cords, separated by very broad concave mid-portion with faint opisthocline riblets. Later teleoconch whorls weakly coronate with 10–14 small, widely spaced, pointed tubercles. Broad concave mid-portion. Suprasutural row of tubercles largely covered by subsequent whorl. Faint spiral sculpture covers entire surface. Suture narrowly incised, weakly undulating around tubercles. Last whorl 67–70% of total height. Subsutural ramp poorly defined, tuberculose. Subsutural ramp deeply concave. Shoulder strongly inflated, delimited by row of small tubercles, concave below to peribasal cord. Peribasal cord with weaker tubercles, strongly constricted at base. Siphonal fasciole poorly delimited, weakly rounded with prominent growth increments; irregular spiral cords of secondary strength at mid-whorl and over base. Aperture pyriform. Outer lip not thickened, smooth within. anal sinus wide, moderately deep, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal long, narrow, slightly deflected in dorsal direction, shallowly notched at tip. Columella strongly excavated in upper half, weakly twisted below, smooth. Columellar callus weakly thickened, forming broad rim; parietal callus thin, adherent.

Discussion. This species is outstanding due to its sculpture of the early teleoconch whorls of very narrow adsutural cords separated by a wide, concave mid-portion. These features distinguish it from all other clavatulid groups. Pleurotoma (Clavatula) clarae Hoernes & Auinger, 1891 was preoccupied by Tenison-Woods (1880: 11) for a Miocene species from Australia. Consequently, Finlay (1927) proposed Clavatula auingeri as replacement name for the Paratethyan species. Finlay (1927) had overlooked that this name was also preoccupied by Pleurotoma (Clavatula) auingeri Hilber, 1879 and by Pleurotoma auingeri Hoernes, 1875a .

‘ Clavatula ’ jarzynkae nov. nom. is reminiscent of ‘ Clavatula’ juliae ( Hoernes & Auinger, 1891) in its outline, but is distinguished by the weaker sculpture and by the different early teleoconch sculpture. In ‘ C. ’ jarzynkae the adsutural cords, are separated by a broad concave mid-portion, whereas in ‘ C. ’ juliae the mid-portion is covered by opisthocline axial riblets that end in a large, swollen beads at the adapical suture. The spire angle of the early whorls is narrower and the spire coeloconoid in ‘ Clavatula’ juliae , whereas the growth rate is constant in ‘ Clavatula ’ jarzynkae giving a regularly conical spire. Moreover, the offshore species ‘ C. ’ juliae seems to have been ecologically separated from the inner neritic ‘ C. ’ jarzynkae.

Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al. 2005; Roetzel 2009). Preservation and reddish color of the fossil shells, suggest, that ‘ Clavatula ’ jarzynkae belonged to the allochthonous shallow water fauna and not to the autochthonous deep-water fauna.

Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin: Korytnica ( Poland) (Friedberg 1928); North Alpine -Carpathian Foreland Basin: Grund ( Austria).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 15 A1–A3. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). B1–B2. ‘Clavatula’ juliae (Hoernes & Auinger 1891), lectotype, NHMW 1949/0005/0043b, Lăpugiu de Sus (Romania). C. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043d, Lăpugiu de Sus (Romania). D1–D3. ‘Clavatula’ jarzynkae nov. nom., NHMW 1949/0005/0043, Grund (Austria). E1–E3. ‘Clavatula’ jarzynkae nov. nom., NHMW 1855/0002/0021, Grund (Austria).