Olegia doderleini ( Hörnes, 1854 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 82-83

Olegia doderleini ( Hörnes, 1854) nov. comb.

Figs 24F 1 –F 3 , G 1 –G 2 , 3 Aa, 5, 7

* Pleurotoma Doderleini Hörn. — Hörnes 1854: 339, pl. 36, figs 17a–b.

Clavatula doderleini (M. Hoernes) — Papp 1954, pl. 9, figs 8–10.

Clavatula doderleini doderleini (M. Hoernes) — Boda 1959: 626, pl. 29, figs 6–7.

C [lavatula]. (C [lavatula]) doderleini (Hörn.) — Sieber 1958a: 156.

Clavatula doderleini doderleini (M. Hoernes) — Papp 1974: 353, pl. 10, figs 12–14.

non Pleurotoma doderleini Hörn. — Pilar 1874: 216, pl. 2, figs 20–21 [= Olegia mandici nov. sp.]. non Pleurotoma Doderleini M. Hoern. — Hoernes 1875b: 69, pl. 2, fig. 4 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula doderleini (Hörn.) — Laskarev 1903: captions of plates, pl. 4, figs 32–34 [= Olegia rumana ( Simionescu & Barbu, 1940)]. non Clavatula Doderleini M. Hoern. — Friedberg 1912: 202, pl. 13, fig. 2 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula Doderleini Hoern. —Friedberg 1928: 571 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Pleurotoma doderleini M. Hörnes — Pauca 1935: 207, pl. 9, figs 3–4 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula Doderleini Hoern. — Friedberg 1938: 143 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Pleurotoma (Clavatula) doderleini Hoern. — Simionescu & Barbu 1940: 125, pl. 2, figs 30–31 [= Olegia curta Boda,

1959]. non Clavatula döderleini [sic] M. Hörns.— Saulea 1956: 80, pl. 3, figs 11–14 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula doderleini doderleini (M. Hoernes, 1856) — Švagrovský 1958: 16, pl. 4, figs 9–11 [= Olegia rumana ( Simionescu & Barbu, 1940)]. non Clavatula (Clavatula) doderleini doderleini (M. Hoernes) —Švagrovský 1959: 223, pl. 9, fig. 3 [= Olegia rumana ( Simionescu & Barbu, 1940)]. non Clavatula doderleini (M. Hoern.) — Švagrovský 1964: 84, pl. 18, fig. 9 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula doderleini (Hoernes) — Roshka 1964: pl. 2, figs 8–9 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula doderleini (M. Hoernes) — Spajić 1966: 44, pl. 4, figs 6–8 [= Olegia rumana ( Simionescu & Barbu, 1940) ]. non Clavatula doderleini (M. Hoernes, 1856) — Zelinskaya et al. 1968: 220, pl. 50, figs 16–17 [= Olegia rumana ( Simionescu & Barbu, 1940)]. non Clavatula (Clavatula) doderleini doderleini (Hörnes, 1856) —Hinculov in Iliescu et al. 1968: 147, pl. 36, figs 11a–b [=

Olegia curta Boda, 1959 ]. non Clavatula (Clavatula) doderleini doderleini (Hoernes, 1856) — Kojumdgieva 1969: 116, pl. 39, figs 4–8 [= Olegia curta Boda, 1959]. non Clavatula doderleini doderleini (M. Hoernes, 1856) — Švagrovský 1971: 398, pl. 70, figs 1–3 [= Olegia rumana ( Simionescu & Barbu, 1940)]. non Clavatula doderleini (Hoern.) — Ali-Zade 1974: 104, pl. 32, fig. 6 [= Nassariidae ].

Type material. Lectotype designated herein: NHMW 1866 /0001/0550, SL: 14.9 mm, MD: 6.5 mm, Wiesen ( Austria), illustrated in Hörnes (1854, pl. 36, figs 17a–b), figs 24F 1 –F 3 . Paralectotypes: NHMW 1858 /0015/0147, SL: 14.6 mm , MD: 6.5 mm, Wiesen ( Austria), figs 24G 1 –G 2, 3 Aa ; NHMW 2021 /0129/0002, 2 spec., Wiesen ( Austria) .

Revised description. Shell small, solid, stout fusiform, with relatively short spire; apical angle ~36–43°. Protoconch not preserved. Teleoconch of nine whorls. Early teleoconch whorls flat-sided, with bipartite sculpture; narrow subsutural cord, suprasutural cord bearing drop-shaped, opisthocline beads. Later teleoconch whorls rapidly increasing in width, with poorly delimited, irregularly beaded subsutural collar and rounded tubercles on suprasutural cord, mid-whorl portion narrow, concave. Suture narrowly and weakly incised, weakly undulating. Last whorl ~65–70% of total height. Subsutural collar broad, bearing large, rounded tubercles. Subsutural ramp narrow, concave, delimited by tubercular shoulder cord, weakly convex below, moderately constricted at base. Siphonal fasciole poorly delimited, weakly rounded. Sculpture of three rows of coarse tubercles.Adapical cords on subsutural collar and at shoulder strongest, and peribasal cord weakest, two further poorly defined and faintly tubercular cords over base. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide, moderately deep, asymmetrically U-shaped, with apex just below collar. Siphonal canal moderately short, straight, shallowly notched at tip. Columella weakly excavated in upper third, straight below, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad callus rim and narrow pseudumbilical chink.

Discussion. Although Hörnes (1854) provided precise illustrations and a clear description of Olegia doderleini , the understanding of this species was distorted by subsequent authors, who erroneously identified and illustrated specimens of O. rumana ( Simionescu & Barbu, 1940) or O. curta ( Boda, 1959) as O. doderleini (see O. rumana for discussion). The short and slightly gradate spire, however, separates O. doderleini clearly from O. rumana , O. curta and O mandici nov. sp.

Overall, O. doderleini is a rare species, known so far only from the middle Sarmatian. An ancestor might be represented by the Badenian O. szokolyensis ( Strausz, 1960) [see discussion for Olegia szokolyensis ( Strausz, 1960) ].

Paleoenvironment. The mollusc assemblages of Nexing and Wiesen suggest costal inner neritic to intertidal environments (Harzhauser & Piller 2010)).

Distribution in Central Paratethys. Sarmatian (middle Miocene): Vienna Basin: Nexing (own data), Eisenstadt-Sopron Basin: Wiesen ( Austria) ( Papp 1974); Pannonian Basin: Budapest, Várpalota, Péccsváradról ( Hungary) ( Boda 1959); Borod Basin: Vârciorog ( Romania) ( Filipescu et al. 2014).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 24 A1–A3. Olegia agathae (Hoernes & Auinger, 1891), lectotype, NHMW 1999z0098/0001, Niederkreuzstetten (Austria). B1–B3. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088a, Niederkreuzstetten (Austria). C. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088b, Niederkreuzstetten (Austria). E1–E2. Olegia curta (Boda 1959), holotype, Sz.1432, Várpalota (Hungary). D1–D2. Olegia szokolyensis (Strausz, 1960), holotype, M.9, Szokolya (Hungary). F1–F3. Olegia doderleini (Hörnes, 1854), lectotype, NHMW 1866/0001/0550, Wiesen (Austria). G1–G2. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria).