Olegia rumana ( Simionescu & Barbu, 1940 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 86-87

Olegia rumana ( Simionescu & Barbu, 1940) nov. comb.

Figs 26E 1 –E 2 , F 1 –F 2 , G 1 –G 2 , 5 , 7

Clavatula doderleini (Hörn.) — Laskarev 1903: captions of plates, pl. 4, figs 32–34 [non Olegia doderleini ( Hörnes, 1854) ]. Clavatula Doderleini M. Hoern. — Friedberg 1912: 202, pl. 13, fig. 2 [non Olegia doderleini ( Hörnes, 1854) ]. Clavatula Doderleini var. striata Friedb. — Friedberg 1912: 203, pl. 13. fig. 3 [non ‘Clavatula’ striata Gray, 1826 ]. Pleurotoma doderleini M. Hörnes — Pauca 1935: 207, pl. 9, figs 3–4. [non Olegia doderleini ( Hörnes, 1854) ]. Pleurotoma (Clavatula) doderleini Hoernes var. sarmatica n.v. — Simionescu & Barbu 1940: 126, pl. 2, figs 32–33. * Pleurotoma rumana n. f.— Simionescu & Barbu 1940: 126, pl. 2, figs 34–35.? Clavatula soceni n. sp. — Jekelius 1944: 87, pl. 23, fig. 16. Clavatula doderleini sarmatica Simionescu et Barbu — Jekelius 1944: 87. Clavatula doderleini doderleini (M. Hoernes, 1856) — Švagrovský 1958: 16, pl. 4, figs 9–11 [non Olegia doderleini ( Hörnes, 1854)]. Clavatula (Clavatula) doderleini rumana (Simionescu et Barbu) — Švagrovský 1958: 17, pl. 5, figs 8–12. Clavatula (Clavatula) soceni Jekelius — Švagrovský 1958: 18, pl. 5, figs 4–7. Clavatula (Clavatula) doderleini striata Friedberg — Švagrovský 1958: 17, pl. 5, figs 1–3 [non ‘Clavatula’ striata Gray, 1826 ]. Clavatula (Clavatula) doderleini doderleini (M. Hoernes) —Švagrovský 1959: 223, pl. 9, fig. 3 [non Olegia doderleini ( Hörnes, 1854)]. Clavatula (Clavatula) doderleini rumana (Simionescu et Barbu) —Švagrovský 1959: 223, pl. 9, fig. 4. Clavatula doderleini (M. Hoern.) — Švagrovský 1964: 84, pl. 18, fig. 9 [non Olegia doderleini ( Hörnes, 1854) ]. Clavatula striata Friedb. — Švagrovský 1964: 84, pl. 18, fig. 10 [non ‘Clavatula’ striata Gray, 1826 ]. Clavatula doderleini (M. Hoernes, 1856) — Zelinskaya et al. 1968: 220, pl. 50, figs 16–17 [non Olegia doderleini ( Hörnes, 1854)]. Clavatula (Clavatula) doderleini sarmatica ( Simionescu & Barbu, 1940) — Kojumdgieva 1969: 117, pl. 39, figs 3. Clavatula doderleini doderleini (M. Hoernes, 1856) — Švagrovský 1971: 398, pl. 70, figs 1–3 [non Olegia doderleini ( Hörnes, 1854)]. Clavatula doderleini striata Friedberg, 1923 — Švagrovský 1971: 401, pl. 70, figs 8–9 [non ‘Clavatula’ striata Gray, 1826 ]. Clavatula rumana ( Simionescu et Barbu, 1940 — Švagrovský 1971: 402, pl. 70, figs 10–14.

Type material. Lectotype designated herein: SL: 17.1 mm, MD: 7.6 mm, Seliște-Orhei ( Moldova), illustrated in Simionescu & Barbu (1940, pl. 2, fig. 34). We are not aware of the whereabouts of the type specimens.

Additional studied material. ZNG PAN A-I- 50/467, SL: 24.5 mm, MD: 9.2 mm, Chlebów ( Poland), illustrated in Friedberg (1912, 203, pl. 13. fig. 2), stored in the Geological Museum of the Institute of Geological Sciences, Polish Academy of Sciences, figs 26E 1 –E 2; ZNG PAN A-I- 50/468, SL: 22.0 mm, MD: 8.2 mm, Chlebów ( Poland), figs 24F 1 –F 2; ZNG PAN A-I- 50/468, SL: 15.0 mm, MD: 6.1 mm, Chlebów ( Poland), figs 26G 1 –G 2; MNG P– 4968, Soceni ( Romania), holotype of Clavatula soceni Jekelius, 1944 , illustrated in Jekelius (1944, pl. 23, fig. 16), stored in the National Museum of Geology of Romania (Bucharest).

Revised description. Small, solid, moderately slender fusiform shell with conical spire; apical angle ~35°. Protoconch not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided with bipartite sculpture; indistinct subsutural cord, weakly concave mid-whorl portion, drop-shaped beaded suprasutural cord. Later whorls with weak subsutural collar bearing rounded to weakly pointed tubercles, concave mid-whorl with comma-shaped riblets, and suprasutural row of larger but weaker tubercles. Suture narrowly incised, weakly undulating, running below abapical row of tubercles. Last whorl ~ 65% of total height; subsutural collar relatively narrow, bearing small, sharp tubercles, subsutural ramp concave, delimited by tubercular shoulder cord, convex below, weakly to moderately constricted at base. Siphonal fasciole poorly delimited, weakly rounded. Sculpture of four narrow, subequal, weakly tubercular primary spiral cords. Adapical cord delimiting shoulder, second cord at mid-whorl delimiting periphery, third peribasal, fourth perifasciolar. Aperture moderately wide, ovoid. Outer lip not thickened, smooth within. anal sinus deep, moderately wide, symmetrically U-shaped, with apex just below collar. Siphonal canal moderately short, strongly deflected to the left, shallowly notched at tip. Columella strongly excavated, twisted at fasciole, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad callus rim.

Discussion. Olegia rumana ( Simionescu & Barbu, 1940) was frequently confused with Olegia doderleini ( Hörnes, 1854) , but differs considerably in its more fusiform shape and higher spire, and in having finer spiral cords on the last whorl below the shoulder bearing much smaller tubercles. This species was first described by Friedberg (1912) as Clavatula Doderleini var. striata , but this name was preoccupied by Gray [1826: 485; = Epideira striata ( Gray, 1826) ] for an extant species from Western Australia. This was overlooked by all subsequent authors, except Tucker (2004: 937). Švagrovský (1971: 401) designated the specimen illustrated by Friedberg (1912, pl. 13, fig. 3) as lectotype. The next available name for this species is Pleurotoma rumana Simionescu & Barbu, 1940 . In the same paper, Simionescu & Barbu (1940) introduced Pleurotoma sarmatica as new species. The type specimen of O. sarmatica is a slender fusiform shell with strongly reduced sculpture. A similar morphotype was described by Kojumdgieva (1969) from the Sarmatian of Bulgaria and by Jekelius (1944) from Romania. In our opinion, Clavatula sarmatica represents specimens of Olegia rumana with reduced sculpture. As first revisers we give C. rumana precedence over C. sarmatica .

Clavatula soceni Jekelius, 1944 is based on a juvenile spire fragment with high conical, paucispiral protoconch and three abapically beaded early teleoconch whorls. The specimen seems to be conspecific with Olegia rumana , and provisionally entered into synonymy.

Paleoenvironment. Coastal inner neritic environments (e.g., Soceni, Lukeneder et al. 2011).

Distribution in Central Paratethys. Sarmatian (middle Miocene): Ukrainian Fore-Carpathian Basin: Chmielnik, Grabki at Szydlów, Chlebów, Dmosice, Dwikozy, Miechocin, Wielowieś, Bogdanówka ( Poland), Wola Mazowiecka, Zalice ( Ukraine) ( Friedberg 1912, 1938); Vienna Basin: Trkmanec (the Czech Republic), Rohožník ( Slovakia) ( Švagrovský 1971); Eisenstadt-Sopron Basin: Nexing ( Austria) (own data), Wiesen ( Austria) ( Papp 1974); Pannonian Basin: Budapest, Várpalota, Péccsváradról ( Hungary) ( Boda 1959); Soceni ( Romania) ( Jekelius 1944); East Slovakian Basin: Bohdanovce, Izra, Kožuchov, Kalša, Kuzmice, Nižný Čaj, Olšovany, Rakoš, Slanec ( Slovakia) ( Švagrovský 1958, 1960a, 1971); Mahida Basin: Domaşnea ( Romania) (Hinculov in Iliescu et al. 1968); Beiuş Basin: Bucuroaia ( Romania); South Carpathian Foreland Basin: Seliște-Orhei, Bihor ( Moldova) ( Simionescu & Barbu 1940); Dacian Basin: Bulgaria ( Kojumdgieva 1969).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 26 A1–A2. Olegia mandici nov. sp., holotype, NHMW 2021/0140/0001, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). B1–B2. Olegia mandici nov. sp., paratype, NHMW 2021/0140/0002, Bekića vrelo, Čremušnica (Croatia). C1–C2. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). D1–D2. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008b, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). E1–E2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/467, Chlebów (Poland). F1–F2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/468, Chlebów (Poland). G1–G2. Olegia rumana (Simionescu & Barbu, 1940), ZNG PAN A-I-50/468, Chlebów (Poland).