Perrona louisae ( Hoernes & Auinger, 1891 )

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 110-111

Perrona louisae ( Hoernes & Auinger, 1891)

Figs 32E 1 –E 3 , F 1 –F 2 , 33E 1 –E 3 , F 1 –F 3 , 5 , 7

Pleurotoma calcarata Grat. — Hörnes 1854: 345, pl. 37, figs 6–9 [non ‘Clavatula’ calcarata Grateloup, 1832 ]. * Pleurotoma (Clavatula) Louisae nobis— Hoernes & Auinger 1891: 354.

Clavatula (Clavatula) louisae Hörnes & Auinger — Glaessner 1926: 116.

C [lavatula]. (C [lavatula]) louisae (R. Hörn. et Au.) — Sieber 1958a: 157.

Clavatula louisae (Hoern. et Auing.) — Kókay 1966: 63, pl. 9, fig. 14.

Perrona louisae (Hörnes & Auinger, 1891) —Harzhauser 2002: 120, pl. 11, figs 8–9.

Type material. Lectotype designated herein: NHMW 1861 /0050/0162a, SL: 25.7 mm, MD: 12.5 mm, Weinsteig ( Austria), illustrated in Hörnes (1854, pl. 37, figs 6a–b), figs 32E 1 –E 3 . Paralectotypes: NHMW 1861 /0050/0162b, SL: 23.2 mm , MD: 11.9 mm, Weinsteig ( Austria) , illustrated in Hörnes (1854, pl. 37, figs 7a–b), figs 32F 1 –F 2; NHMW 1861 /0050/0162c, SL: 20.6 mm , MD: 9.8 mm, Weinsteig ( Austria) , illustrated in Hörnes (1854, pl. 37, figs 8a–b), figs 33E 1 –E 3; NHMW 1859 /0045/0010a, ( Austria) , illustrated in Hörnes (1854, pl. 37, figs 9a–b); NHMW 1861 /0050/0162, SL: 27.6 mm , MD: 11.7 mm, Weinsteig ( Austria) , figs 33F 1 –F 3; NHMW 1861 /0050/0162, 28 spec., Weinsteig ( Austria) ; NHMW 1859 /0045/0010, 19 spec., Weinsteig ( Austria) ; 27 spec., NHMW 1846 /0037/0313, Weinsteig ( Austria) .

Additional studied material. NHMW2000 z0001/0049, Stetten ( Austria) , illustrated in Harzhauser (2002, pl. 11, figs 8–9); NHMW 1861 /0050/0087, 41 spec., Niederkreuzstetten ( Austria) ; NHMW 1861 /0050/0027, 5 spec., Rückersdorf ( Austria) , NHMW 1865 /0001/1089, 1 spec., Grund ( Austria) ; NHMW 2007 z0071/0038, 5 spec., Gainfarn ( Austria) .

Revised description. Shell medium-sized, solid, stocky, with strongly coronate and gradate spire and low cylindrical teleoconch whorls; apical angle ~50°. Protoconch not preserved. Teleoconch of at least seven whorls. Early teleoconch whorls with smooth, prominent, rounded subsutural cord, concave below, covered with fine narrow spiral cords separated by narrow interspaces; weak comma-shaped riblets on early teleoconch whorls. Later teleoconch whorls squat, cylindrical, subsutural cord strengthens to form collar with large, rounded to pointed tubercles, of variable strength and often poorly defined. Suture narrowly but moderately deeply incised, weakly undulating. Last whorl ~ 65% of total height barrel-shaped; subsutural collar strong, coronate, bearing 10–12 tubercles, delimited by weak groove in some specimens. Subsutural ramp broad, shallow, weakly concave, delimited by weak, low-set shoulder cord, placed close below peribasal cord. Base weakly angled, but more so than shoulder. Base moderately constricted, concave. Peribasal cord weakly developed, siphonal fasciole weak, slightly twisted. No further spiral sculpture. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within. Anal sinus deep, moderately wide, asymmetrically U-shaped, with apex just below subsutural collar. Siphonal canal moderately long, wide, slightly deflected, shallowly notched at tip. Columella strongly excavated in adapical half, slightly twisted at fasciole, smooth. Columellar and parietal callus slightly thickened, sharply delimited, forming broad callus rim.

Discussion. Perrona lousiae ( Hoernes & Auinger, 1891) is characterized by its broad, strongly gradate and strongly coronate spire, squat barrel-shaped whorls, and the absence of spiral sculpture on later adult whorls, apart from the major cords. It resembles Perrona descendens ( Hilber, 1879) in general shape, but the latter has a narrower spire angle, higher spire whorls and lacks pointed tubercles. The same features distinguish P. lousiae from Perrona vindobonensis (Quenstedt, 1884) , Perrona ilonae nov. sp. and P. sabinae ( Hoernes & Auinger, 1891) .

Paleoenvironment. The occurrence at Weinsteig suggests a preference for lagoonal environments with mudflats ( Harzhauser et al. 2002; own data).

Distribution in Central Paratethys. Karpatian (early Miocene): Korneuburg Basin: Grossrussbach, Kleinebersdorf, Karnabrunn, Rückersdorf, Wetzleinsdorf, Weinsteig ( Austria) (Harzhauser 2002); North AlpineCarpathian Foreland Basin: Niederkreuzstetten ( Austria) (Harzhauser 2002, 2003); Badenian (middle Miocene): North Alpine Foreland Basin: Grund ( Austria) ( Hoernes & Auinger 1891); Vienna Basin: Gainfarn ( Austria) (hoc opus); Pannonian Basin: Herend ( Hungary) ( Kókay 1966).

Occurrences from the Ottnangian (early Miocene) of Kaltenbachgraben in Germany, reported by Steininger (1973) will need confirmation.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 32 A1–A3. Perrona loetschi nov. sp., holotype, NHMW 1949/0005/0102a, Grund (Austria). B1–B2. Perrona loetschi nov. sp., paralectotype, NHMW 1949/0005/0102b, Grund (Austria). C1–C3. Perrona loetschi nov. sp., paralectotype, NHMW 2021/0153/0001, Grund (Austria). D1–D2. Perrona loetschi nov. sp., paralectotype, NHMW 1855/0045/0752, Grund (Austria). E1–E3. Perrona louisae (Hoernes & Auinger, 1891), lectotype, NHMW 1861/0050/0162a, Weinsteig (Austria). F1–F2. Perrona louisae (Hoernes & Auinger, 1891), paralectotype, NHMW1861/0050/0162b, Weinsteig (Austria).

FIGURE 33 A1–A3. Perrona sabinae (Hoernes & Auinger 1891), lectotype, NHMW 1949/0005/0104a, Lăpugiu de Sus (Romania). B1–B3. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0104b, Lăpugiu de Sus (Romania). C1–C3. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 1854/0035/0249, Lăpugiu de Sus (Romania). D1–D3. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). E1–E3. Perrona louisae (Hoernes & Auinger, 1891), paralectotype, NHMW1861/0050/0162c, Weinsteig (Austria). F1–F2. Perrona louisae (Hoernes & Auinger, 1891), paralectotype, NHMW1861/0050/0162, Weinsteig (Austria).