Perrona barbarae ( Hoernes & Auinger, 1891 ), 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 120-121

publication ID	https://doi.org/10.11646/zootaxa.5123.1.1
publication LSID	lsid:zoobank.org:pub:036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A
DOI	https://doi.org/10.5281/zenodo.10722952
persistent identifier	https://treatment.plazi.org/id/039487D1-FFF5-FFD0-FFBA-FC786B7CFDD9
treatment provided by	Plazi (2022-03-30 20:52:14, last updated by Valdenar 2024-02-29 15:27:11)
scientific name	Perrona barbarae ( Hoernes & Auinger, 1891 )
status	comb. nov.

Treatment

Perrona barbarae ( Hoernes & Auinger, 1891) View in CoL nov. comb.

Figs 36E View FIGURE 36 1 –E View FIGURE 1 3 View FIGURE 3 , F 1 –F View FIGURE 1 3 View FIGURE 3 , 4L View FIGURE 4 , 5 View FIGURE 5 , 7 View FIGURE 7

Clavatula Barbarae —Hoernes 1891: 131 [nomen nudum].

* Pleurotoma (Clavatula) Barbarae nov. form.— Hoernes & Auinger 1891: 355, pl. 48, figs 12–13.

C [lavatula]. (C [lavatula]) barbarae (R. Hörn. et Au.) View in CoL — Sieber 1958a: 157.

Clavatula barbarae (Hörnes & Auinger, 1891) —Harzhauser 2002: 118, pl. 10, figs 14–15.

Type material. Lectotype designated herein: NHMW 1949 View Materials /0005/0103a, Rückersdorf ( Austria), SL: 12.8 mm, MD: 5.2 mm, illustrated in Hoernes & Auinger (1891, pl. 48, figs 13a–b) and Harzhauser (2002, pl. 10, fig. 14), figs 36E 1 –E 3 . Paralectotypes: NHMW 1949 View Materials /0005/0103b, Rückersdorf ( Austria), SL: 12.8 mm , MD: 5.0 mm, illustrated in Hoernes & Auinger (1891, pl. 48, figs 12a–b) and Harzhauser (2002, pl. 10, fig. 14), figs 36F 1 –F 3; NHMW 1976 View Materials /1785/0035, Kleinebersdorf ( Austria), illustrated in Harzhauser (2002, pl. 10, fig. 15) .

Additional studied material. NHMW 1861 View Materials /0050/0027, 1 spec., Rückersdorf ( Austria) .

Revised description. Shell small, solid, moderately broad fusiform, with gradate spire; apical angle ~43°. Protoconch and earliest teleoconch whorls not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided with narrow subsutural spiral cord and sinuous, strongly opisthocline, comma-shaped axial riblets ending in weak beads at abapical suture. Profile of later whorls distinctly concave. On fourth whorl subsutural cord developing into swollen subsutural collar, tubercular on last two whorls. Axial sculpture fading leaving smooth suprasutural cord. Suture narrowly impressed, weakly undulating. Last whorl 65% of total height, subcylindrical. Subsutural collar broad, rounded, bearing large, wide-set, indistinct rounded tubercles. Subsutural ramp broad, concave, delimited by rounded, indistinctly tubercular shoulder cord. Base concave delimited by slightly narrower, smooth, rounded perifasciolar cord. Siphonal fasciole rounded, slightly twisted with weak growth increments. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately deep and wide, asymmetrically U-shaped, with apex just below subsutural collar. Siphonal canal moderately long, narrow, twisted, slightly deflected to the left, moderately notched at tip. Columella strongly excavated, twisted at fasciole, smooth. Columellar and parietal callus thickened, adherent, forming broad callus rim.

Discussion. This species was treated so far in Clavatula , but the opisthocline axial riblets on the early teleoconch whorls suggest placement in Perrona . Perrona variocingulata ( Sacco, 1890) , from the Tortonian of Stazzano, S. Agata and Montegibbio ( Italy) ( Sacco 1904, pl. 12, figs 61, 62; Montanaro 1937, pl. 5, figs 31–34) is highly reminiscent of Perrona barbarae ( Hoernes & Auinger, 1891) . The illustrated specimens differ from P. barbarae in the slightly lower position of the beads on the subsutural collar, the less concave last whorl, the weaker basal and peribasal spiral cords, and the presence of secondary spiral cords between primary spiral cords and on the base. Similarly, Perrona agassizi ( Bellardi, 1847) , from the Tortonian of Italy, has a subcylindrical whorl profile, pointed beads in a low position on the subsutural collar and develops beads on the shoulder (see Ferrero Mortara et al. 1981, pl. 14, figs 9a–b).

Perrona variocingulata sensu Martelli & Nelli, 1910 , from the middle Miocene of Albania, is clearly distinguished from P. barbarae View in CoL by its granulose spiral sculpture (see Martelli & Nelli 1910: 549, pl. 11, fig. 3).

Paleoenvironment. Estuarine, littoral mud flats ( Zuschin et al. 2004).

Distribution in Central Paratethys. Karpatian(early Miocene): Korneuburg Basin:Rückersdorf,Kleinebersdorf ( Austria) (Harzhauser, 2002, 2003, Zuschin et al. 2004).

References

Bellardi, L. (1847) Monografia delle Pleurotome fossili del Piemont. Stamperia Reale, Torino, 122 pp. [https: // www. biodiversitylibrary. org / item / 120573 # page / 3 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 58977

Ferrero Mortara, E., Montefameglio, L., Pavia, G. & Tampieri, R. (1981) Catalogo dei tipi e degli esemplari figurati della collezione Bellardi e Sacco. Parte I. Museo Regionale di Scienze Naturali, Catalogo, 6, 1 - 327. [https: // www. biodiversitylibrary. org / item / 269998 # page / 5 / mode / 1 up]

Hoernes, R. & Auinger, M. (1891) Die Gasteropoden der Meeres-Ablagerungen der ersten und zweiten Miocanen Mediterran- Stufe in der Osterreichisch-Ungarischen Monarchie. Abhandlungen der kaiserlich-koniglichen Geologischen Reichsanstalt, 12, 283 - 382, pls 37 - 50. [https: // opac. geologie. ac. at / ais 312 / dokumente / Hoernes % 20 und % 20 Auinger % 20 _ 1879 _ Gasteropoden. pdf]

Martelli, A. & Nelli, B. (1910) Miocene medio e superiore di Valone in Albania. Bollettino della SocietaI geologica italiana, 29, 513 - 551. [https: // www. biodiversitylibrary. org / item / 209913 # page / 697 / mode / 1 up]

Montanaro, E. (1937) Studi monografici sulla malacologia miocenica modenese. Parte I. I mollusci tortoniani di Montegibbio. Palaeontographia Italica, New Series 7, 37, 115 - 191.

Sacco, F. (1890) Catalogo paleontologico del bacino terziaro del Piemonte. Bollettino della Societa Geologica Italiana, 9, 189 - 340. [https: // www. biodiversitylibrary. org / item / 210076 # page / 207 / mode / 1 up]

Sacco, F. (1904) I molluschi dei terreni terziari del Piemonte e della Liguria. Parte 30. Aggiunte e correzioni (con 1400 figure). Considerazioni generali. Indice generale dell'opera. C. Clausen, Torino, 203 + xxxvi pp., 31 pls. [https: // www. biodiversitylibrary. org / item / 46270 # page / 90 / mode / 1 up]

Sieber, R. (1958 a) Systematische Ubersicht der jungtertiaren Gastropoden des Wiener Beckens. Annalen des Naturhistorischen Museums in Wien, 62, 123 - 192. [https: // opac. geologie. ac. at / ais 312 / dokumente / Annalen _ des _ NHMW _ 1958 _ 062 _ 123 _ 192. pdf]

Zuschin, M., Harzhauser, M. & Mandic O. (2004) Palaeoecology and taphonomy of a single parautochthonous Paratethyan tidal flat deposit (Karpatian, Lower Miocene - Kleinebersdorf, Lower Austria). Courier Forschungsinstitut Senckenberg, 246, 153 - 168.

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 36 A1–A3. Perrona emmae (Hoernes & Auinger 1891), lectotype, NHMW 1949/0005/0113a, Enzesfeld (Austria). B1–B3. Perrona emmae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0113b, Enzesfeld (Austria). C1–C3. Perrona emmae (Hoernes & Auinger 1891), paralectotype, NHMW 2013/0479/1523, Gainfarn (Austria). D1–D3. Perrona emmae (Hoernes & Auinger 1891), NHMW 2013/0479/1523, Gainfarn (Austria). E1–E3. Perrona barbarae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0103a, Rückersdorf (Austria). F1–F3. Perrona barbarae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0103b, Rückersdorf (Austria).