Perrona barbarae ( Hoernes & Auinger, 1891 ), 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 120-121

Perrona barbarae ( Hoernes & Auinger, 1891) nov. comb.

Figs 36E 1 –E 3, F 1 –F 3, 4L, 5, 7

Clavatula Barbarae —Hoernes 1891: 131 [nomen nudum].

* Pleurotoma (Clavatula) Barbarae nov. form.— Hoernes & Auinger 1891: 355, pl. 48, figs 12–13.

C [lavatula]. (C [lavatula]) barbarae (R. Hörn. et Au.) — Sieber 1958a: 157.

Clavatula barbarae (Hörnes & Auinger, 1891) —Harzhauser 2002: 118, pl. 10, figs 14–15.

Type material. Lectotype designated herein: NHMW 1949 /0005/0103a, Rückersdorf ( Austria), SL: 12.8 mm, MD: 5.2 mm, illustrated in Hoernes & Auinger (1891, pl. 48, figs 13a–b) and Harzhauser (2002, pl. 10, fig. 14), figs 36E 1 –E 3 . Paralectotypes: NHMW 1949 /0005/0103b, Rückersdorf ( Austria), SL: 12.8 mm , MD: 5.0 mm, illustrated in Hoernes & Auinger (1891, pl. 48, figs 12a–b) and Harzhauser (2002, pl. 10, fig. 14), figs 36F 1 –F 3; NHMW 1976 /1785/0035, Kleinebersdorf ( Austria), illustrated in Harzhauser (2002, pl. 10, fig. 15) .

Additional studied material. NHMW 1861 /0050/0027, 1 spec., Rückersdorf ( Austria) .

Revised description. Shell small, solid, moderately broad fusiform, with gradate spire; apical angle ~43°. Protoconch and earliest teleoconch whorls not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided with narrow subsutural spiral cord and sinuous, strongly opisthocline, comma-shaped axial riblets ending in weak beads at abapical suture. Profile of later whorls distinctly concave. On fourth whorl subsutural cord developing into swollen subsutural collar, tubercular on last two whorls. Axial sculpture fading leaving smooth suprasutural cord. Suture narrowly impressed, weakly undulating. Last whorl 65% of total height, subcylindrical. Subsutural collar broad, rounded, bearing large, wide-set, indistinct rounded tubercles. Subsutural ramp broad, concave, delimited by rounded, indistinctly tubercular shoulder cord. Base concave delimited by slightly narrower, smooth, rounded perifasciolar cord. Siphonal fasciole rounded, slightly twisted with weak growth increments. Aperture moderately narrow, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately deep and wide, asymmetrically U-shaped, with apex just below subsutural collar. Siphonal canal moderately long, narrow, twisted, slightly deflected to the left, moderately notched at tip. Columella strongly excavated, twisted at fasciole, smooth. Columellar and parietal callus thickened, adherent, forming broad callus rim.

Discussion. This species was treated so far in Clavatula , but the opisthocline axial riblets on the early teleoconch whorls suggest placement in Perrona . Perrona variocingulata ( Sacco, 1890) , from the Tortonian of Stazzano, S. Agata and Montegibbio ( Italy) ( Sacco 1904, pl. 12, figs 61, 62; Montanaro 1937, pl. 5, figs 31–34) is highly reminiscent of Perrona barbarae ( Hoernes & Auinger, 1891) . The illustrated specimens differ from P. barbarae in the slightly lower position of the beads on the subsutural collar, the less concave last whorl, the weaker basal and peribasal spiral cords, and the presence of secondary spiral cords between primary spiral cords and on the base. Similarly, Perrona agassizi ( Bellardi, 1847) , from the Tortonian of Italy, has a subcylindrical whorl profile, pointed beads in a low position on the subsutural collar and develops beads on the shoulder (see Ferrero Mortara et al. 1981, pl. 14, figs 9a–b).

Perrona variocingulata sensu Martelli & Nelli, 1910 , from the middle Miocene of Albania, is clearly distinguished from P. barbarae by its granulose spiral sculpture (see Martelli & Nelli 1910: 549, pl. 11, fig. 3).

Paleoenvironment. Estuarine, littoral mud flats ( Zuschin et al. 2004).

Distribution in Central Paratethys. Karpatian(early Miocene): Korneuburg Basin:Rückersdorf,Kleinebersdorf ( Austria) (Harzhauser, 2002, 2003, Zuschin et al. 2004).