Lloydiella japenensis, Ballantyne & Lambkin, 2009

Ballantyne, Lesley A. & Lambkin, Christine, 2009, Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae), Zootaxa 1997, pp. 1-188 : 63-64

Lloydiella japenensis sp. n.

( Figs 205, 209, 212–214, 217–219, 222)

Holotype. Male. INDONESIA IRIAN JAYA: 1.45S, 136.15E, Japen Is, Mt Baduri , 1000 ft, VIII. 1938 L Cheesman ( NHML).

Paratypes (3). Same data as for holotype, female, 2 males ( NHML) .

Diagnosis. One of a group of three species that strongly resemble Ll. majuscula in colour, size, and shape, distinguished as follows: Ll. uberia sp. n. has rounded posterolateral corners on the pronotum (the other three have angulate corners); Ll. majuscula is Australian, and has a well–defined longitudinal groove on the ventral side of T8 (the other three species lack such a development); Ll. wareo sp. n. has lateral margins that converge posteriorly, posterolateral corners subequal to 90º, and antennal sockets almost contiguous ( Ll. japenensis sp. n. has lateral pronotal margins that diverge along their length, posterolateral corners less than 90º, and antennal sockets are not contiguous).

Male. 10.5–10.6 mm long; 4.3–4.4 mm wide; W/L 0.4. Colour ( Figs 205, 209): Pronotum orange, semitransparent, fat body visible in irregular clumps; MS and MN slightly paler; elytra very dark brown; head, antennae, palpi, apical ¼ of femora and tibiae and tarsi of all legs, abdominal V2–5, and tergites 2–6, almost black; ventral aspect of thorax pale yellow; tergites 7, 8 pale yellow. Pronotum: 2.0– 2.1 mm long; 3.5 mm wide; W/L 1.6; lateral margins diverge along their length with very slight convergence in 1/3; posterolateral corners angulate, <90° obliquely inclined to median line and projecting beyond median posterior margin. Elytron: with 2 well–defined interstitial lines (1, 2) and fainter traces of line 3. Head: GHW 2.1–2.3 mm; SIW 0.3; SIW/GHW 0.1; ASD<ASW (sockets close but not contiguous). Apical segment of labial palpi with 2 teeth (developed at base and apex of inner longer edge), basal tooth in 1/ 3 males with a slight emargination (appears as 2 very short teeth). Abdomen, ventrites ( Fig. 222): LO extends to sides and close to posterior margin of V7. MPP short broad and curves slightly dorsally along its length (may be a postmortem effect). Tergites: T8 quite well sclerotised; wider than long in posterior visible area and lateral margins converge slightly posteriorly; ventral surface lacking median longitudinal groove. Aedeagus Figs 213–215. Aedeagal sheath ( Figs 218–220) with sheath sternite very broad in posterior half.

Etymology. The specific name is Latinised and genitive case from the locality name.