Trichotichnus (Bottchrus) schawalleri

Schmidt, Joachim, 2017, Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species, Zootaxa 4323 (3), pp. 301-358: 315-318

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Trichotichnus (Bottchrus) schawalleri

sp. n.

Trichotichnus (Bottchrus) schawalleri  sp. n.

( Figs 38View FIGURES 38 – 41, 42–46View FIGURES 42 – 46, 151View FIGURE 151, 5View FIGURES 5 – 13)

Type material. Holotype: ♂, “539 NEPAL: Bhojpur Distr. Dilkharka 2100 m, 26.V.1997 leg. W. SCHAWALLER”, “Mus. Stuttg” ( SMNS). 

Paratype: Nepal: 1 ♂, “536 NEPAL: Bhojpur Distr. Valley NW Phedi 1900 m, 25.V.1997 leg. W. SCHAWALLER”, “Mus. Stuttg” (cJS).

Description (male). Habitus as in Fig. 38View FIGURES 38 – 41. Body length in holotype 5.8, in paratype 6.1 mm, width 2.7 and 2.8 mm, respectively.

Colour: Body piceous, shiny on dorsum, with light bluish or blue greenish tinge; labrum, very narrow margins of pronotum and apical portion of elytral margins as well as of elytral epipleurae slightly paler; palpi, antennae and legs brownish yellow.

Head: Comparatively large, slightly smaller than in T. (B.) bubsaensis  sp. n., with wide neck (HWmax/PWmax = 0.67, HWmin/PWmax = 0.55–0.57). Eyes rather convex (HWmax/HWmin = 1.17–1.22), but slightly less than in T. (B.) bubsaensis  sp. n., in lateral view elongate oval. Tempora about half as long as eye, slightly convex, somewhat abruptly sloped to neck. Genae almost twice as wide as antennomere 1, with few short setae. Lateral margin of head between antenna and eye, like in all the preceding species, slightly protruded laterally, forming an obtuse angle near eyes, very finely bordered. Area between supraorbital furrow and upper margin of eye wide, approximately as wide as width of antennomere 2 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum slightly concave anteriorly. Clypeus very slightly emarginate along anterior margin, moderately convex basally, flatly sloped to apex, with a comparatively large setigerous pore at each anterior corner. Frons and vertex convex. Mentum and submentum completely fused, submentum with one long seta and a shorter one laterally of the long seta on each side. Left mandible not truncate, slightly blunted at apex. Dorsal microsculpture consisting of obliterate weakly transverse meshes. Antennae (in male) almost extending to the basal edge of pronotum, with antennomeres 4–8 markedly longer than wide.

Pronotum: Comparatively wide (PWmax/PL = 1.41), widest at the end of anterior third, moderately narrowed posteriad (PWmax/PWmin = 1.18–1.19), with one lateral seta inserted slightly before the widest point. Sides rounded almost throughout, slightly sinuate just before basal angles. Apical margin very slightly emarginate, bordered only laterally. Apical angles very slightly prominent anteriad, widely rounded at tip. Basal margin nearly straight, only slightly roundly oblique laterally, vaguely bordered in lateral third, slightly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles slightly more than 90, each with a sharp, subdenticulate apex slightly protruded laterad. Pronotal disc moderately convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally although latero-basal areas slightly flattened. Lateral gutter very narrow, very hardly widened in apical third. Median line interrupted, very fine, superficial, ended markedly before apical and basal margins. Lateral depressions absent; basal foveae small, somewhat elongate and very shallow. Surface sparsely, somewhat finely and irregularly punctate mainly in latero-basal areas, with middle basal portion impunctate, punctation at apical margin absent. Microsculpture present throughout, distinct; meshes mostly transverse, between apical margin and anterior transverse impression more or less isodiametric.

Elytra: In lateral and caudal view convex, in dorsal view oval, relatively wide (in male, EL/EW = 1.30–1.31; EL/PL = 2.32–2.45; EW/PWmax = 1.27–1.32), widest approximately at middle, with somewhat acute apex. Humeri prominent, rather widely angularly rounded at tip, without denticle. Subapical sinuation shallow. Sutural angle acute and somewhat sharp at tip. Basal edge markedly sinuate laterally, forming distinct obtuse angle with lateral margin. Striae fine, almost superficial along entire length. Intervals somewhat flat throughout, weakly narrowed posteriad. Parascutellar setigerous pore present, large. Parascutellar striole rudimentary, almost absent. Interval 3 with a very small discal setigerous pore at stria 2 behind middle. Marginal umbilicate series widely interrupted at middle, with 6–8 pores in anterior group and 8–9 pores in posterior group. Microsculpture fine, poorly recognizable, consisting of thin transverse lines.

Hindwings: Reduced to tiny scales.

Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum ( Fig. 42View FIGURES 42 – 46) markedly wider than long, strongly narrowed posteriad. Apex of last visible (VII) abdominal sternite (in male) subtruncate and slightly concave just at tip ( Fig. 43View FIGURES 42 – 46).

Legs: Protibia without longitudinal sulcus on upper surface. Profemur with four setae on anterio-ventral margin. Tarsi short, similar to those in T. (B.) parvulus  sp. n., metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) markedly widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two pairs of ventro-lateral setae. In male, pro- and mesotarsi moderately widened.

Male genitalia: Median lobe of aedeagus ( Figs 44–46View FIGURES 42 – 46) in lateral view rather evenly arcuate, C-shaped, with almost straight middle portion of ventral margin and with apical third curved ventrad; in dorsal view almost straight, narrowed to apex in distal third. Terminal lamella moderately short, narrowed to apex. Apical capitulum in lateral view ( Fig. 44View FIGURES 42 – 46) very oblique, poorly delimited, almost not protruded ventrad and dorsad, narrowly rounded at tip; in caudal view ( Fig. 45View FIGURES 42 – 46) triangular, somewhat horseshoe-shaped, rather acute, only slightly blunted at tip. Internal sac without distinct spiny patches and other sclerotic elements.

Etymology. This new species is named after the famous coleopterologist Wolfgang Schawaller (Stuttgart), who collected the type specimens.

Comparison. Trichotichnus (B.) schawalleri  sp. n. is very similar to T. (B.) holzschuhi  and all the other preceding species in habitus and in having metepisternum short, protibia not sulcate dorsally, and aedeagal median lobe arcuate, with inclined apical capitulum, but easily recognized from them in the combination of the following characters: pronotum rather wide, moderately narrowed basad; dorsal microsculpture present throughout on head, pronotum and elytra; parascutellar striole rudimentary, almost absent; apex of last visible (VII) abdominal sternite in male ( Fig. 43View FIGURES 42 – 46) subtruncate and slightly concave just at tip [in this character, T. (B.) schawalleri  sp. n. is similar only to T. (B.) hingstoni  ]; median lobe ( Figs 44–46View FIGURES 42 – 46) with shorter terminal lamella and very oblique, horseshoeshaped apical capitulum, which is poorly separated from the terminal lamella in lateral view and almost triangular [as in T. (B.) hingstoni  ], with rather acute, only slightly blunted tip, in caudal view ( Fig. 45View FIGURES 42 – 46); and internal sac with characteristic pattern of spiny folders in the middle portion of the median lobe. Trichotichnus (B.) cyanescens  , probably occurring sympatricly with T. (B.) schawalleri  sp. n., is somewhat similar to the new species in habitus, but easily distinguished by having larger size (6.1–6.9 mm), metepisternum ( Fig. 115View FIGURES 115 – 119) slightly more elongate, mentum and submentum fused only laterally, apex of last (VII) abdominal sternite ( Fig. 116View FIGURES 115 – 119) in male rounded, and median lobe of aedeagus ( Figs 117–119View FIGURES 115 – 119) S-shaped, hooked at apex. Trichotichnus (B.) hingstoni  , also occurring sympatricly with this new species, is easily distinguished from in having larger size (6.8–7.7 mm), genae glabrous, metepisterna ( Fig. 137View FIGURES 137 – 145) markedly longer, pronotum never sinuate basally and with narrow, apically not widened lateral gutter, protibia longitudinally sulcate on dorsal side, and apical capitulum of the aedeagal median lobe ( Figs 135–136View FIGURES 135 – 136, 140View FIGURES 137 – 145) more strongly prominent dorsally and ventrally.

Distribution ( Fig. 151View FIGURE 151, 5View FIGURES 5 – 13). Up to today only known from the western slope of the middle Arun River Valley towards Salpa pass, Bhojpur District, East Nepal, at altitudes of approximately 1900–2100 m. This species seems to occur sympatricly with T. (B.) hingstoni  ( Fig. 152View FIGURE 152, 6View FIGURES 5 – 13). Another species, T. (B.) cyanescens  , was found in the Arun River Valley slightly further north ( Fig. 151View FIGURE 151, 12View FIGURES 5 – 13).


Staatliches Museum fuer Naturkund Stuttgart