Mesopodagrionidae Kalkman & Abbott, 2021

Bybee, Seth M., Kalkman, Vincent J., Erickson, Robert J., Frandsen, Paul B., Breinholt, Jesse W., Suvorov, Anton, Dijkstra, Klaas-Douwe B., Cordero-Rivera, Adolfo, Skevington, Jeffrey H., Abbott, John C., Herrera, Melissa Sanchez, Lemmon, Alan R., Lemmon, Emily Moriarty & Ware, Jessica L., 2021, Phylogeny and classification of Odonata using targeted genomics, Molecular Phylogenetics and Evolution 160, pp. 107115-107115 : 10

Mesopodagrionidae Kalkman & Abbott fam.n.

(type genus: Mesopodagrion McLachlan, 1896 )

— fairly large (hindwing 27–33 mm) and sturdy damselflies restricted to streams in southern China and the north of Vietnam, Thailand and Myanmar. Wings clear, with two Ax and numerous intercalated veins distally in radial fields; arculus roughly at three-fifth of distance between wing base and node; quadrangle without cross-veins; R4 originates clearly proximal of subnode; IR3 originates at subnode. Pterostigma rectangular, about three times as long as broad; yellow-reddish or black. Adult perches with wings outstretched and abdomen held in horizontal position. Head, thorax, legs and abdomen black with yellowish or blue pattern, including antehumeral stripes that cross the humeral suture to continue onto the mesepimeron and (in males) the pale dorsum of abdominal segments 9 and 10. The postocular lobes are swollen and emphasized by the largely pale (yellow) back of the head, but lack postocular spots. Adult male cerci with a simple forcipate shape, paraprocts short, about a fifth the length of the cerci. Hind rim of abdominal segment 10 modified medially with two short but sturdy spines directed distally. Genital ligula with terminal lobe reduced, deeply incised, and with two long and slender horns; its shaft with <10 setae on each side which are shorter than half the width of the shaft. Nymph is stocky and easily recognized by the large, flat and fanlike horizontal gills (only shared with the unrelated Argiolestidae and Protolestidae ), as well as the occipital lobes that protrude distinctly at the side of the head and are are covered densely with strong spines. Included genera: Mesopodagrion .