Aleocharina Fleming, 1821

Asenjo, Angélico, Irmler, Ulrich, Klimaszewski, Jan, Chandler, Donald S., Fierros-López, Hugo Eduardo & Vieira, Juliana S., 2019, Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution, Zootaxa 4621 (1), pp. 1-406 : 89-92

Subtribe Aleocharina Fleming, 1821

0295. Aleochara Gravenhorst, 1802: 67 ( Figs. 41–42 )

Type species: Staphylinus curtulus Goeze, 1777

Correa Fauvel, 1878c: 592 [synonymized by Pace 2003b: 124]

Type species: Correa oxytelina Fauvel, 1878

Fauvelia Tate, 1880 : xlvi [synonymized by Blackwelder 1952: 165]

Type species: Correa oxytelina Fauvel, 1878

Fauvelia Tate in Holder, 1882: 78

Type species: Correa oxytelina Fauvel, 1878

Subgenera:

Aidochara Casey, 1906: 145

Type species: Aidochara planiventris Casey, 1906

Aleochara Gravenhorst, 1802: 67 [*]

Type species: Staphylinus curtulus Goeze, 1777

Copiata Gozis, 1886: 12 [synonymized by Bernhauer & Scheerpeltz 1 926: 775]

Type species: Staphylinus curtulus Goeze, 1777

Baryodma Thomson, 1858: 31

Type species: Staphylinus bipunctatus Olivier, 1795

Arybodma Blackwelder, 1952: 63 [ Schülke & Smetana 2015: 492]

Type species: Aleochara intricata Mannerheim, 1830

Calochara Casey, 1906: 149 [*]

Type species: Calochara rubripennis Casey, 1906

Oreochara Casey, 1906: 148 [synonymized by Klimaszewski 1984: 59]

Type species: Oreochara laramiensis Casey, 1906

Ceranota Stephens, 1839: 351

Type species: Aleochara daltoni Stephens, 1832

Ceronota Agassiz, 1846: 72 [ Newton & Thayer 2003]

Type species: Aleochara daltoni Stephens, 1832

Hoplonotus Schmidt-Göbel, 1846: 245 [synonymized by Märkel 1846: 300]

Type species: Hoplonotus laminatus Schmidt-Göbel, 1846

Hylozyras Iablokoff-Khnzorian, 1960: 1882 [synonymized by Assing 2015a: 16]

Type species: Zyras stenocephalus Iablokoff-Khnzorian, 1960

Sorecocephala Bernhauer, 1902: 238 [synonymized by Smetana in Löbl & Smetana 2004: 33]

Type species: Ocalea reitteri Bernhauer, 1900

Coprochara Mulsant & Rey, 1874b: 430 [*]

Type species: Aleochara bilineata Gyllenhal, 1810

Eucharina Casey, 1906: 165 [synonymized by Blackwelder 1952: 166]

Type species: Aleochara sulcicollis Mannerheim, 1843

Funda Blackwelder, 1952: 166 [synonymized by Klimaszewski 1984: 15]

Type species: Aleochara sulcicollis Mannerheim, 1843

Mecorhopalus Solier, 1849: 347 [synonymized by Bernhauer & Scheerpeltz 1 926: 775]

Type species: Mecorhopalus elongatus Solier, 1849

Skenochara Bernhauer & Scheerpeltz, 1926: 795 [synonymized by Maus 1 998: 83]

Type species: Aleochara squalithorax Sharp, 1888

Echochara Casey, 1906: 176

Type species: Rheochara lucifuga Casey, 1893

Rheocharella Casey, 1906: 181 [synonymized by Klimaszewski 1984: 94]

Type species: Aleochara fenyesi Bernhauer, 1905

Emplenota Casey, 1884a: 17 [*]

Type species: Emplenota maritima Casey, 1884

Polistoma Casey, 1893: 289 [synonymized by Bernhauer & Scheerpeltz 1926: 795]

Type species: Aleochara obscurella Gravenhorst, 1806

Polycharina Reitter, 1909: 28 [synonymized by Fenyes 1921b: 415]

Type species: Aleochara grisea Kraatz, 1856

Polystoma Stephens, 1833b: 91 [synonymized by Casey 1893: 289]

Type species: Aleochara obscurella Gravenhorst, 1806

Polystomaria Reitter, 1909: 28 [synonymized by Scheerpeltz 1925: 447]

Type species: Aleochara obscurella Gravenhorst, 1806 Heterochara Mulsant & Rey, 1874b: 299

Type species: Aleochara clavicornis Redtenbacher, 1849

Aphaenochara Maruyama & Hlaváč, 2003: 225 [synonymized by Tronquet 2009: 96]

Type species: Aphaenochara lackneri Maruyama & Hlaváč, 2003

Ctenochara Casey, 1906: 134 [ Newton & Thayer 2003]

Type species: Aleochara clavicornis Redtenbacher, 1848

Homoeochara Mulsant & Rey, 1874b: 414

Type species: Aleochara sparsa Heer, 1839

Madagaschara Klimaszewski et al., 2000: 250

Type species: Aleochara subsulcata Pace, 1999

Maseochara Sharp, 1883: 154 [*]

Type species: Maseochara opacella Sharp, 1883

Pinalochara Casey, 1906: 177 [synonymized by Klimaszewski 1984: 103]

Type species: Pinalochara wickhami Casey, 1906

Tithanis Casey, 1884a: 16 [synonymized by Bernhauer & Scheerpeltz 1926: 797]

Type species: Aleochara valida LeConte, 1858

Mesochara Blackwelder, 1952: 239

Type species: Aleochara almorensis Cameron, 1939

Polystomota Casey, 1906: 136

Type species: Aleochara grisea Kraatz, 1856

Tinotus Sharp, 1883: 170 [*]

Type species: Tinotus cavicollis Sharp, 1883

Acrimea Casey, 1911: 14 [synonymized by Gusarov 2003e: 10]

Type species: Acrimea resecta Casey, 1911

Arpatheta Blackwelder, 1952: 62 [synonymized by Sawada 1987: 148]

Type species: Paratheta carnivora Cameron, 1920

Exaleochara Keys, 1907: 102 [synonymized by Bernhauer & Scheerpeltz 1926: 713]

Type species: Aleochara morion Gravenhorst, 1802

Paratheta Cameron, 1920a: 269 [synonymized by Blackwelder 1952: 62]

Type species: Paratheta carnivora Cameron, 1920

Triochara Bernhauer, 1901c: 373

Type species: Aleochara trisulcata Weise, 1877

Xenochara Mulsant & Rey, 1874b: 344 [*]

Type species: Aleochara decorata Aubé, 1850

Dyschara Mulsant & Rey, 1874b: 425 [synonymized by Assing 2009b: 36]

Type species: Aleochara inconspicua Aubé, 1850

Euryodma Reitter, 1909: 24 [synonymized by Assing, 2009b: 36]

Type species: Aleochara brevipennis Gravenhorst, 1806

Isochara Bernhauer, 1901a: 440 [ Klimaszewski 1984: 35]

Type species: Aleochara tristis Gravenhorst, 1806

Metalea Mulsant & Rey, 1875e: 299 [synonymized by Bernhauer & Scheerpeltz 1926: 789]

Type species: Ocalea procera Erichson, 1839

Notiochara Casey, 1906: 134 [synonymized by Klimaszewski & Jansen 1993: 68]

Type species: Notiochara subaspera Casey, 1906

Ophiochara Bernhauer, 1901a: 439 [synonymized by Assing 2009b: 36]

Type species: Aleochara breiti Ganglbauer, 1897

Polychara Mulsant & Rey, 1874b: 348 [synonymized by Klimaszewski 1984: 34]

Type species: Aleochara discipennis Mulsant & Rey, 1853

Rheochara Mulsant & Rey, 1874b: 294 [synonymized by Assing 2016a: 270]

Type species: Ocalea spadicea Erichson, 1837

97 species are included in this genus from Latin America.

Characters/Description: Lynch-Arribálzaga 1884: 67, 74 (characters of subgenus Baryodma ); Fenyes 1920: 389 (characters as Fauvelia ), 396 (characters); Fenyes 1921b: 417 (characters of subgenus Maseochara ); Klimaszewski 1984: 7, 14 (characters of subgenus Coprochara ), 24 (characters of subgenus Xenochara ), 59 (characters of subgenus Calochara ), 70 (characters of subgenus Aleochara ), 89 (characters of subgenus Echochara ), 95 (characters of subgenus Emplenota ), 102 (characters of subgenus Maseochara ); Lohse et al. 1990: 125 (characters in key), 137 (characters); Klimaszewski & Maus 1999: 209 (characters of Puberula species group), 211 (characters of Weiseri species group), 212 (characters of Richteri species group), 215 (characters of Lacustris species group), 216 (characters of Glabra species group), 217 (characters of subgenus Aleochara ), 217 (characters of Parvicollis species group), 218 (characters of Bruchi species group), 219 (characters of subgenus Maseochara ), 219 (characters of Wickhami species group); Hanley 2002b: 456 (characters as Tinotus ); Ashe 2002b: 124 (characters in key), 125 (characters in key of subgenus Tinotus ); Klimaszewski et al. 2002: 284 (characters of subgenus Tinotus ); Gusarov 2003e: 10 (characters of subgenus Tinotus ); Pace 2008a: 364 (characters in key of subgenus Tinotus ); Aballay et al. 2014: 103 (characters in key); Yamamoto & Maruyama 2016: 83 (characters of subgenus Tinotus ).

Key: Lynch-Arribálzaga 1884: 67 (species of Buenos Aires province, Argentina); Fenyes 1920: 397 (some subgenera); Klimaszewski 1984: 14 (subgenera), 16 (species of the subgenus Coprochara ), 36 (species of the subgenus Xenochara ), 60 (species of the subgenus Calochara ), 71 (species of the subgenus Aleochara ), 90 (species of the subgenus Echochara ), 96 (species of the subgenus Emplenota ), 103 (species of the subgenus Maseochara ); Klimaszewski & Ashe 1992: 400 (some species of Costa Rica); Gusarov 2003e: 11 (species of the subgenus Tinotus ).

Distribution.

West Indies: Cuba, Jamaica, Haiti, Puerto Rico, US Virgin Isl., Guadeloupe, Dominica, St. Lucia, St. Vincent & Grenadines, Grenada, Trinidad-Tobago.

Central America: Mexico, Guatemala, Honduras, El Salvador, Nicaragua, Costa Rica, Panama.

South America: Colombia, Venezuela, Suriname, French Guiana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Uruguay, Chile, Argentina.

Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental, Australian, Oceanic.

0296. Ocyota Sharp, 1883: 163

Type species: Ocyota dubia Sharp, 1883

2 species are known from and are restricted to Central America. Assing & Wunderle (1997: 122), removed Ocyota bruchi Bernhauer (described of Argentina) from the genus but did not place it in any other genus.

Characters/Description: Fenyes 1920: 362 (characters); Assing & Wunderle 1997: 119 (characters); Ashe 2002b: 124 (characters in key).

Distribution.

Central America: Mexico, Guatemala.

Figures

FIGURES 40–51. Habitus; Habrocerinae (40), Aleocharinae (41–51); 40, after Solier 1849; 41, Brazil: state of Paraná (MUSM); 42, Chile: region of Antofagasta (MUSM); 43, after Pace 2015e; 44, after Ashe 2002a; 45, after Pace 2014a; 46, after Pace 2012; 47, after Y.H. Kim & Ahn 2015; 48, after Pace 2014c; 49, Chile: region of Araucanía (MUSM); 50, after Irmler 2011; 51, after Caron & Ribeiro-Costa 2008; slightly modified. Scale bars: 1 mm.