Thraulodes flavus

Kluge, Nikita J., 2020, Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species, Zootaxa 4756 (1), pp. 1-142 : 83

Group flavus

Definition. In imago and subimago of both sexes, thorax light, with interrupted blackish stripe along pleura; fore wing maculate, costal field with many cross veins bordered by brown, pterostigmatic cross veins anastomozed; femora twice banded; abdominal terga II–VI of male with oblique lateral streaks, without midway spots ( Figs 365–367 , 398–401 ). Hind wing without dark brown coloration of costal brace ( Figs 376–408 ) (in contrast to all examined species).

Composition. Th. flavus sp. n. and Th. spangleri Traver & Edmunds 1967 .

Figures

FIGURES 365–370. Thraulodes flavus sp. n. 365–366, male imago in lateral and dorsal view; 367, female imago; 368, left half of subimaginal exuviae of mesonotum; 369, subimaginal exuviae of fore leg of male; 370, genitalia of male imago (368–370, holotype).

FIGURES 396–402. Thraulodes spangleri. 396–397, left half of subimaginal exuviae of mesonotum of two different specimens; 398–400, male imago in lateral, dorsal and ventral view; 401, female imago; 402, genitalia of male imago.

FIGURES 371–376. Thraulodes flavus sp. n. 371, genitalia of male imago in lateral view; 372, apex of penis in dorsal view; 373, genitalia of male imago in ventral view; 374, their subimaginal exuviae; 375, larval protogonostyli and protopenis; 376, hind wing (372–376, holotype).

FIGURES 377–378. Thraulodes flavus sp. n., eggs.

FIGURES 379–388. Thraulodes spangleri. 379–383, larval exuviae at the same magnification: 379–381, parts of head with labrum and mandibles; 382, right half of pronotum and mesonotum; 383, abdominal terga and sterna; 384, abdominal terga and sterna of male subimago extracted from larva; 385, tergalius; 386, larval fore tibia with subimaginal tibia inside and hypodermal dark brown markings; 387, anterior side of larval hind femur; 388, anterior side of larval hind tibia.

FIGURES 389–395. Thraulodes spangleri, labrum. 389–391, one specimen: 389, general view with focus on dorsal side and posterior transverse setal row; 390, focus on ventral side and denticles; 391, outline of denticles separated by four sensilla (shown as dots). 392–395, another specimen: 392, focus on dorsal side and posterior setal row; 393, focus on anterior transverse setal row (transformed to field of irregularly situated setae); 394, focus on denticles and tips of irregularly situated setae of the anterior row; 395, outlines of vestigial denticles separated by incisions and two sensilla.

FIGURES 403–408. Thraulodes spangleri. 403–404, genitalia of male imago in lateral and ventral view; 405, apex of penis in dorsal view; 406, subimaginal exuviae of styliger, gonostylus and spear-like rolls; 407, larval protogonostyli and protopenis; 408, hind wing.