Fidia longipes (Melsheimer)

Strother, M. S. & Staines, C. L., 2008, A revision of the New World genus Fidia Baly 1863 (Coleoptera: Chrysomelidae: Eumolpinae: Adoxini), Zootaxa 1798 (1), pp. 1-100 : 45-49

Fidia longipes (Melsheimer)

( Figs. 14 , 46 , 93 ; Map 3)

Eumolpus longipes Melsheimer 1847: 169 (original description).

Melsheimer (1847) did not state the number of specimens he had before him when he described E. longipes . However, he provided a single length measurement, "2 l[ines].” (= approximately 4.23 mm.), for this species, whereas he provided ranges of length measurements for many other new species throughout the publication. This would suggest that Melsheimer's description was based on a single specimen, and no evidence was found to contradict this assumption.

In a letter dated April 28, 1875, to Alexander Agassiz, J. L. LeConte indicated that he had all of the unique types of the Melsheimer collection ( Anonymous 1961). Five specimens fitting the description of E. longipes are housed in the LeConte collection of the MCZC. One female labeled as originating in Illinois, "Ill.", is actually a specimen of F. rileyorum , n. sp. Melsheimer described E. longipes from Pennsylvania and clearly this was not the specimen before Melsheimer. Among the four remaining specimens, are three females bearing the following labels: "[pink disc] / P. viticolus ! Uhler [handwritten] / J.L. LeConte Collection / Fidia longipes (Melsheimer) det. M.S.Strother 1993"; "[pink disc] / J.L. LeConte Collection / Fidia longipes (Melsheimer) det. M.S.Strother 1993"; and "J.L. LeConte Collection / Fidia ♀ nr. longipes det. M.S.Strother 1993". The females measure 4.48, 5.04, and 4.98 mm., respectively and clearly do not agree with the measurement of approximately 4.23 mm. given by Melsheimer. The fourth specimen is a male which bears the following labels: "longipes!Mels. [handwritten] / J.L. LeConte Collection / HOLOTYPE Eumolpus longipes Melsheimer 1847 [red, added by M. S. Strother]". The male measures 4.20 mm. and matches Melsheimer's description in all other respects as well is here taken to be the Holotype. The holotype is in good condition, pinned through the right elytron at the suture, and missing the following: left eleventh antennomere, right antennomeres 6–11, and the right metathoracic leg. The aedeagus was not dissected.

The Melsheimer collection at the MCZC contains five specimens (2 ♂♂, 3♀♀) bearing the labels, " Pennsylvania Melsheimer / Melsheimer Collection / Fidia longipes (Melsheimer) det. M.S.Strother 1993". Schultz (1970) designated one of the males, which he dissected, as the lectotype but never validly published this designation .

Pachnephorus viticolus Uhler 1855: 418 (original description); Crotch 1873: 34 (as synonym of F. longipes ).

The type was not found and is assumed to be lost. Designation of a neotype was not deemed necessary in this case to maintain nomenclatural stability. Horn & Kahle (1935 –37) stated that various parts of Uhler's collection were located in the MCZC, USNM, and CASC, respectively, but they made no mention of Uhler's Coleoptera collection. The MCZC does not contain a specimen labeled as Uhler's type (M. S. Kelley pers. comm.), and the specimen in the LeConte collection bearing the handwritten label, " P. viticolus ! Uhler", is much smaller than the length stated in the description. No reply was received regarding the Uhler specimens in the CASC. Staines was unable to locate any specimens in the USNM labeled as Uhler’s type. It was hoped that his types might be located in the ANSP because

Uhler published his description in the Proceedings of the Academy of Natural Sciences of Philadelphia; however, no Uhler types were found in this collection (D. Azuma pers. comm.) .

Based on the type locality of Baltimore, Maryland, Uhler's name can only apply to two taxa, F. longipes and F. viticida . Although the length measurement of 2½ l[ines]. (= approximately 5.29 mm.) given by Uhler is slightly larger than the largest specimen of F. longipes examined during this revision (5.16 mm.), the description of the leg coloration clearly indicates that the taxon Uhler had before him was F. longipes .

Pachnephorus (Fidia) viticolus: Walsh 1867a: 88 .

Pachnephorus (Fidia) longipes: Walsh 1867a: 88 .

Fidia longipes: Walsh 1867b: 118 ; Crotch 1873: 34 (faunal treatment); Henshaw 1885: 108 (checklist); Lefèvre 1885: 75 (catalog); Horn 1892: 198,199 (key, diagnosis); Schaeffer 1904: 228 (key); Clavareau 1914: 76 (catalog); Leng 1920: 293 (catalog); Fattig 1948: 19 (faunal treatment); Wilcox 1954: 402 (faunal treatment); Schultz 1970: 245 (dissertation); Balsbaugh & Hays 1972: 73–74 (faunal treatment); Wilcox 1975: 57 (checklist); Downie & Arnett 1996: 1334 (key); Riley et al. 2003: 151 (catalog); Clark et al. 2004: 103 (host plants); Ciegler 2007: 169 (faunal treatment).

Fidia cana: Fattig 1948: 19 (misidentification, faunal treatment).

Description. Males: TL = 3.72–5.04 mm, HW = 1.72–2.20 mm. Females: TL = 4.08–5.16 mm, HW = 1.96– 2.76 mm. Color: Extremely variable; dorsum fulvous to black, most lighter specimens often with at least some dark coloration on head, lateral aspect of pronotum, and/or sutural and lateral elytral margins; thoracic and abdominal sterna fulvous to black; femora, especially meso- and metathoracic pairs, almost always basally lighter (some specimens from throughout range rarely with all femora entirely red-brown), tibiae and tarsi fulvous to black; pubescence whitish to golden-yellow. Pronotum: Length subequal to width, widest at middle, sides moderately to distinctly arcuate in dorsal view, dorsum gently convex to nearly straight in lateral view; densely, coarsely punctate-reticulate; pubescence moderately sparse to moderately dense, not obscuring surface sculpture. Mesepisternum: Densely pubescent. Elytra: Intrahumeral callus obsolescent to feebly developed; asetose punctate-striae well-developed, occasionally obscured by pubescence; interstices flat to convex, moderately densely to densely punctulate-strigulose to punctulate-undulose; pubescence moderately sparse to dense, usually appearing uniformly distributed, occasionally arranged in feeble to moderately distinct longitudinal rows between striae. Abdomen: Males: medial area of first three sterna feebly flattened; first sternum, and occasionally second and third as well, with small impunctate, glabrous area; pygidium dorsally gently convex in apical ½ with broadly rounded apex. Females: medial area of all sterna evenly convex, uniformly punctate-pubescent; apical margin of last sternum feebly concave with or without small, rounded, poorly developed medial process; pygidium dorsally with shallow, transversely lunate preapical impression; apex slightly reflexed, acutely rounded to subtruncate. Legs: Both sexes with femora gradually tapered towards base. Male with protibia distally slightly curved ventrally; apex slightly dorsoventrally compressed and laterally expanded, with ventral surface shallowly excavated; pro- and mesotibial spurs stout, minutely rugulose; protibial spurs separated by at least the length of one spur, mesotibial spurs contiguous; disco-setae on pro- and mesobasitarsi only. Penis: In posterior view, sides feebly concave to nearly straight, widest at bases of apical lobes; apical emargination small, width slightly greater than depth, basally with small medial tooth usually well-developed. In lateral view, eudorsal surface feebly to moderately convex; euventral surface feebly to moderately concave; distally tapered to acute apex. Sperm guide composed of lower sclerite only. Spermatheca: Basal arm type.

Diagnosis. Small to medium-sized (3.72–5.16 mm.); fulvous to black, with at least one pair of femora bicolorous, distinctly lighter at base than in distal portion (rarely with all femora unicolorous); pronotal punctation coarse. Male with protibia slightly dorsoventrally compressed and laterally expanded at apex; protibial spurs separated by at least the length of one spur ( Fig. 46 ); apical emargination of penis small, width slightly greater than depth, with small medial tooth usually well-developed ( Fig. 93 ).

Fidia longipes is very similar to F. delilahae and F. rileyorum . Characters distinguishing F. delilahae from F. longipes were given in the diagnosis for the former species. Fidia longipes and F. rileyorum are very similar, and females of the two species cannot be distinguished without associated males. Males of F. longipes

have the protibial spurs separated by at least the length of one spur, whereas males of F. rileyorum have the protibial spurs separated by little more than the basal width of one spur. In addition, the apical emargination of the penis is wider and deeper in F. rileyorum than it is in F. longipes . Finally, the two species are largely allopatric, with sympatry occurring only along the Appalachian Mountains, as well as in central Ohio in the North and central Alabama in the South.

Distribution (Map 3). Fidia longipes occurs predominantly east of the Appalachian Mountains from southern New York west to central Ohio south to northern Florida and central Alabama. Specimens were collected at elevations ranging from near sea level to approximately 1220 m.

Specimens examined (696). UNITED STATES. ALABAMA: Autauga Co., 1 mi. S Pine Level, v:20:1990 (RHT:3); Barbour Co., 4 mi. W. of Hwy 431 on Hwy 82, v:31:1992 (MSS:23); Bullock Co., Union Springs, vi:29:1965 (AUEM:1); Cleburne Co., 2.3 mi. S. of Hwy 49 on Hwy 281, vi:1:1992 (MSS:3); Coffee Co., Ft. Rucker Mil. Res., vi:26:1983 (FSCA:1, RHT:1); Dale Co., Ft. Rucker Mil. Res., vi:11:1983 (EGRC:1, RHT:4), vi:12:1983 (FSCA:10, RHT:4); Elmore Co., hwy 231, 6.2 mi. S jct. hwy 14, v:28:1989 (RHT:4), vi:4:1989 (RHT:5); Henry Co., 3.2 mi. N. of Hwy 431 on Hwy 173, v:31:1992 (MSS:2); Houston Co., at the jct. of Hwy 231 and 109, v:31:1992 (MSS:1); Lee Co., 1.3 mi. N. of Hwy 80 on Hwy 51, v:31:1992 (MSS:8), Auburn, vii:1959 (AUEM:1); Macon Co., 3 mi. E of Y, vi:11:1963 (AUEM:1), hwy 80, 1 mi. E jct. Co. Rd. 49, vi:4:1989 (RHT:2); Russell Co., 3 mi. N. of county line on Hwy 51, v:31:1992 (MSS:7), 5.1 mi. S. of Lee Co. line on Hwy 51, v:31:1992 (MSS:5); Shelby Co., nr. Helena, v:18:1957 (FMNH:3); Tallapoose Co., 1 mi. NW Reeltown, vii:31:1964 (AUEM:1), 6 mi. N. of Hwy 280 on Hwy 49, v:31:1992 (MSS:1), Horseshoe Bend Nat'l Pk., vi:25:1964 (AUEM:1). DISTRICT OF COLUMBIA: Washington, vi:3:1902 (USNM:1), vi:4:1903 (USNM:1), vi:6 (USNM:2), vi:14:1903 (USNM:2), vi:19:1903 (USNM:3), vii:29:1907 (USNM:3), vi:17:1909 (USNM:1), vi:19:1909 (USNM:1), vi:19:1911 (USNM:4), vii:1:1923 (AUEM:1), vi:93 (USNM:1), no date (MCZC:2, DEFW:3, USNM:2), Washington; Rock Creek Pk., vi:8:1907 (USNM:1). FLORIDA: Calhoun Co., Clarksville at Chipola R., v:22:1986 (CMNC:1); Gadsen Co., 6 mi. N Quincy; C- 161 Willacoochee Cr., v:25:1988 (FAMU:1), Rocky Comfort Cr.; 5.3 mi SW Quincy, vi:14:1988 (FAMU:1); Jackson Co., 4 mi. S of Marianna, v:26:1965 (NDSU:2), 4.5 mi. N Altha, v:14:1983 (FSCA:1), FL Caverns St. Pk., v:19:1985 (EGRC:21, LSUC:12); Liberty Co., Torreya St. Pk., v:18:1970 (FSCA:1), v:3:1976 (AMNH:3, EGRC:3), v:4:1976 (AMNH:1, EGRC:1), vi:6:1982 (FSCA:2), vii:9:1988 (RHT:1), vi:6:1982 (RHT:1), v:7:1989 (RHT:1), vii:8:1989 (RHT:1), Torreya State Pk., v:19:1985 (EGRC:9, LSUC:1); Madison Co., 2.7 mi. E. of Jefferson Co. line, v:26:1992 (MSS:1), 7.9 mi. E. of the Jefferson Co. line, v:26:1992 (MSS:1); Suwannee Co., 3.0 mi. E. of I-10 on Hwy 90, v:26:1992 (MSS:1); County not determined: River Junction, v:16:1939 (FSCA:6). GEORGIA: Bartow Co., Stamp Creek; Weems-Macbeth Property, vi:29:1990 (FSCA:1); Carroll Co., Vila Rica, vi:17:42 (UGCA:1); Chattahoochee Co., 2.5 mi. W of Marion Co. line on Hwy 26, v:25:1992 (MSS:7); Clarke Co., Whitehall Forest, vii:14:1976 (UGCA:1), vi:25–30:1976 (UGCA:1), vi:6–9:1978 (RHT:1); Dekalb Co., Stone Mt., v:29:36 (UGCA:1); Gilmer Co., Ellijay, vi:27:31 (UGCA:1); Habersham Co., Cornelia, vi:22:1906 (CUIC:1); Hall Co., 4 mi. NW Lula, vi:14:1980 (RHT:3, UGCA:1); Harris Co., 4 mi. W. of FDR St. Pk. on Hwy 90, v:25:1992 (MSS:7); Jackson Co., Hardeman Forest, vii:1–6:1974 (UGCA:1), vii:21–28:1974 (UGCA:1), viii:1974 (UGCA:1); Jefferson Co., jct. hwy 242 & 171, vi:17:1978 (RHT:1); Liberty Co., St. Catherine's Is., v:27:1978 (CSUC:17); Marion Co., 6.1 mi. W. of Schley Co. line on Hwy 26, v:25:1992 (MSS:1); Murray Co., Fort Mt., vii:9:37 (UGCA:1); Oconee Co., 2 mi. NW Watkinsville; nr. jct. 53 & 207, vii:12:1981 (UGCA:1); Rabun Co., Clayton, vii:1910 (AMNH:1), no date (UAIC:2); Thomas Co., 0.4 mi. N. of FL border on Hwy 19, v:26:1992 (MSS:7), 4.4 mi. W. of Hwy 19 on Hwy 84, v:26:1992 (MSS:7); Towns Co., Appalachian Trail at hwy 75, vii:4:1986 (RHT:3), hwy 180, 1– 3 mi. W jct. hwy 75, vii:11:1987 (RHT:1); State only, no date (FSCA:1, USNM:1). MARYLAND: Anne Arundel Co., Odenton, vi:5:1918 (CUIC:3, USNM:2), vii:4:1918 (CUIC:1); Baltimore, vi:24:09 (CHSC:32), vi:29:09 (CASC:16), vi:30:09 (CASC:3), vii:2:09 (BPBM:1), vii:15:09 (CASC:1), vii:21:09 (CASC:1), vii:2:1940 (USNM:1), no date (BMNH:2), Baltimore Co., Catonsville, vi:21:37 (CNCI:3), North Point, vi:22:1940 (USNM:1), Sparrows Point, vii:5:34 (CASC:9), vii:5:34 (FMNH:3); Frederick Co., county only, no date (MCZC:3); Montgomery Co., Glen Echo, Su/1922 (USNM:9), vi:28:50 (USNM:2), Great Falls, vii:2:05 (USNM:1), viii:2:05 (USNM:1), vi:17:1913 (USNM:1), vii:12:18 (USNM:2), vi:25:1963 (CASC:1), Plummers I, vi:27:05 (USNM:3), vii:4:07 (USNM:2), vii:7:07 (USNM:1), viii:7:07 (USNM:1), vi:20:09 (USNM:1), Plummers Id, vi:17:1913 (INHS:1), county only, vi:17:11 (USNM:1), vi:25:11 (USNM:1); Prince George’s Co., Beltsville, vii:4:15 (USNM:1), Bladensburg, vi:13:16 (USNM:1); State only, no date (CUIC:2, MCZC:2, DEFW:5). NEW JERSEY: State only, no date (USNM:1). NEW YORK: Orange Co., West Point, ix:15:1912 (USNM:1); State only, no date (AMNH:1). NORTH CAROLINA: Buncombe Co., Black Mt., vii:4:40 (CASC:3), Blue Ridge Pkway, vi:25:1973 (JEWC:1); Haywood Co., Lake Junaluska, vi:20–23:1977 (FSCA:2), nr. Canton Rt 23,vi:18:1978 (NCSU:2), near Canton; Cody Vineyard, vi:28:1978 (NCSU:1), nr Fines Creek, vi:14–15:1984 (FSCA:1); Johnston Co., nr Clayton; Poole Vineyard, vi:8:1978 (NCSU:2), v:28:1979 (NCSU:4), vi:4:1979 (NCSU:1), vii:14:1979 (NCSU:1); Macon Co., Coweeta Hydrologic Lab, vii:3:1983 (RHT:2); Madison Co., Hot Springs, no date (AMNH:2); Moore Co., nr. West End; Auman Vineyard, vi:15:1978 (NCSU:12); Onslow Co., near Jacksonville; Melville Farms, vi:23:1978 (NCSU:1); Orange Co., Chapel Hill, vi:34 (UCDC:1); Perquimans Co., nr. Hertford; Perry Vineyard, vi:24:1978 (NCSU:6); Rutherford Co., nr. Bostic; Newton Vineyard, vi:18;1978 (NCSU:1); Stanly Co., county only, vii:23:59 (NCSU:1); Wake Co., Raleigh, v:31:1952 (CMNC:1); Warren Co., W of Norlina; road 158; field W-42, viii:15:1979 (NCSU:1); Yadkin Co., SW of Marler; road 1116; field Y-8, vii:6:1979 (NCSU:1); County not determined: Cherry, vi:3:1959 (NCSU:1), Southern Pines, vi:7:1952 (CMNC:1), vi:14:1952 (CMNC:2); State only, no date (MCZC:1, USNM:1). OHIO: Athens Co., Athens, vi:24:35 (UGCA:1); Clinton Co., county only, vi:19:62 (OSUC:1); Delaware Co., Hwy 745, Eversole Run, vii:5:1986 (BYUC:3), jct. Deer Creek & Scioto River, vii:21:1984 (BYUC:1), county only, vi:21:50 (OSUC:1); Franklin Co., Columbus, vii:13:62 (OSUC:1), Columbus, Mock Park, vii:14:1984 (BYUC:2), county only, vii:11:42 (CDAE:1), vii:24:48 (OSUC:1); Highland Co., county only, vii:17:58 (FSCA:2), viii:13:60 (FSCA:2), vii:16:61 (FSCA:4); Hocking Co., county only, vi:19:38 (OSUC:1); Knox Co., Gambler, no date (FSCA:1); Lake Co., Willoughby, vi:26:1901 (OSUC:2); Meigs Co., Bedford Tunp., vii:28:39 (UGCA:2); Scioto Co., county only, vi:17:44 (OSUC:1); Vinton Co., Lake Hope, vi:28:1987 (BYUC:1), vii:12:1987 (BYUC:3), Lake Hope State Park, vi:20:1984 (BYUC:4), Vinton Furnace near Dundas, vi:19:1985 (BYUC:1). PENNSYLVANIA: Allegheny Co., Allegheny, vii:2:94 (CUIC:4), vii:10:97 (CUIC:4), no date (INHS:1, LACM:6, MCZC:3, NDSU:1, PURC:3); Berks Co., S.C.M., vii:13:57 (CNCI:1); Cumberland Co., Mt. Holly Spgs., vii:11:1920 (LACM:1), New Cumb., vii:7:40 (NDSU:1); Dauphin Co., Harrisburg, vii:14 (INHS:1), viii:19:1975 (NDSU:1), Hummelstn, vii:4:18 (ANSP:1), Paxtang, vii:26:62 (NDSU:2), Stoverdale, vi:16:1964 (AUEM:1), county only, viii:15:27 (CASC:1); Delaware Co., Chadds Ford, vii:15–30:1987 (JEWC:1), vi:19:1988 (FSCA:1), vii:8:1989 (JEWC:3), vii:4:1990 (JEWC:4), Darby, vii:4:00 (FMNH:1), Glenolden, vi:16 (USNM:1), Swarthmore, vi:13 (OSUC:1); Franklin Co., 3 mi. W Roxbury, vii:15:1989 (JEWC:1); Indiana Co., Angora, vi (MCZC:4); Montgomery Co., Abington, vi (MCZC:4), Glenside, vi:17:06 (USNM:1); Philadelphia Co., Frankford, no date (USNM:2), Philadelphia, vi:22:99 (USNM:1), vi:23:06 (USNM:1), no date (MCZC:1), W Park, no date (OSUC:3); Westmoreland Co., Jeanette, vi:1 (NCSU:1), vi:8 (AMNH:1), vi:11 (AMNH:1); County not determined: Clark's Valley, vii:7:40 (NDSU:1); State only, no date (UAIC:1). SOUTH CARO- LINA: Aiken Co., Aiken , v:30:57 (CMNC:1), v:31:57 (CMNC:1, CNCI:2), vi:22:1957 (CNCI:1); Greenville Co., county only, vii:24:1935 (CUCC:1); Pickens Co., Clemson College, vii:30:1926 (CUCC:2), vi:13:1927 (CUCC:1), vii:2:1928 (CUCC:1), vii:12:1932 (CASC:1); County not determined: Rocky Bottom, viii:12:1958 (NDSU:1). TENNESSEE: Anderson Co., Oak Ridge, no date (CDAE:1); Johnson Co., Johnson C., vi:12:1951 (CASC:1); Monroe Co., 11 mi. W Vonore, vi:12:1955 (CMNC:1), Vonore, vi:12:1955 (FSCA:1); Sevier Co., Great Smoky Mts. N.P.; Elkmont, viii–ix:27-1:1986 (MUIC:1). VIRGINIA: Arlington Co., Arlington, vii:4:19 (USNM:1), vii:31:19 (USNM:1); Botetourt Co., Craig Creek Rec. A nr. Oriskany, vi:14:1991 (EGRC:3); Fairfax Co., Great Falls, vi:19:10 (USNM:1), no date (MCZC:1), Gt. Falls, vi:22:12 (USNM:2), vi:10:16 (USNM:2), county only, vi:23:20 (USNM:1); Lee Co., Pennington Gap, no date (MCZC:2, USNM:3); Loudoun Co., Bluemont, vi:28:1914 (USNM:1); Nelson Co., county only, vii:1:1913 (USNM:2), vi:26:1914 (USNM:2), vii:4:1915 (USNM:7); Orange Co., county only, vii:19 (USNM:1); Spotsylvania Co., Fredrkbg, v:12:00 (USNM:1), vi:19:04 (USNM:4); independent city, Alexan., vi:26:1911 (USNM:1); County not determined: 4-Mile Run, vi:29:1913 (USNM:1), Acquia Cr., v:24:96 (ANSP:1, MCZC:1), Glencarlyn, vi:20:1917 (CUIC:1), Rosslyn, vii:4 (USNM:1), viii:07 (USNM:3), Shenandoah Pk. & Blue Ridge, vii:23:51 (CASC:1); State only, no date (AMNH:1). WEST VIRGINIA: Berkeley Co., Allensville, vii:6:1977 (BYUC:1); Greenbriar Co., W. Sulphur, vii:3:1912 (USNM:1), vii:11:1913 (USNM:1), vii:18:1914 (USNM:2), White Sulphur, vii (CASC:2); Kanawha Co., Guthrie, vii:16:1989 (BYUC:1), viii:5:1989 (BYUC:2), vii:10:1990 (BYUC:1), vii:21:1990 (BYUC:1), viii:24:1990 (BYUC:1), viii:29:1990 (BYUC:1), vii:2:1991 (BYUC:1), vi:4:1991 (BYUC:6), Kanawha St. For., vi:16:1987 (BYUC:1), vi:19:1987 (BYUC:1); Lincoln Co., Hamlin, vii:26:1968 (BYUC:1); Mason Co., McClintic Wildl. Sta., vii:7:1980 (BYUC:1), vii:11:1980 (BYUC:1), vii:19:1980 (BYUC:2), viii:6:1980 (BYUC:1), vii:27:1980 (BYUC:4), McClintic Wildlife Sta., viii:21:1991 (BYUC:2); Mercer Co., Bluefield, vi:29:1989 (BYUC:1); Monroe Co., Hollywood, viii:23:1979 (BYUC:1), viii:29:1979 (BYUC:1); Tyler Co., Sistersville, vi:18:30 (CUIC:1); Wirt Co., Burn Spgs, no date (MCZC:1); State only, no date (MCZC:1). No locality, no date (ANSP:6).

Temporal Data. Collecting dates ranged from 3 May to 15 September.

Natural History. Plant associations included Ilex opaca Ait. (Aquifoliaceae) , soybeans ( Fabaceae ), Salix (Salicaceae) , and various Vitaceae including Ampelopsis arborea (L.), Parthenocissus quinquefolia (L.), and various Vitis spp. Specimens were also collected at lights, by "Lindgren funnel", and in Malaise and sticky traps. McGiffen & Neunzig (1985) discussed the seasonal abundance of F. longipes in North Carolina vineyards and illustrated feeding damage caused by this species.

Taxonomic History. This species was first described as Eumolpus longipes by Melsheimer in 1847. Uhler (1855) described the taxon as a new species from Baltimore under the name Pachnephorus viticolus . Walsh (1867a) used Melsheimer's longipes in combination with the generic name Pachnephorus (Fidia) and in a subsequent article ( Walsh 1867b) combined longipes with the generic name Fidia , and this combination has been followed by all subsequent authors.

Fattig (1948) listed F. cana Schaeffer as having been collected by W. T. Davis in June in Clayton, Georgia. It is highly unlikely, if not impossible, that this is an accurate record for F. cana , which has only been collected in central and southwest Texas. The specimen(s) mentioned by Fattig was not seen, although a specimen of F. longipes collected by W. T. Davis in July in Clayton is housed in the AMNH. Fattig did not specify whether Clayton was the county in the central western part of the state or the town in the extreme northeastern notch of the state, but because the other localities listed by Fattig for F. longipes and F. viticida are towns or specific localities rather than counties, it is assumed that the latter is correct. Only F. longipes and F. viticida are known to occur in northeastern Georgia, but the smaller, darker F. longipes more closely matches Schaeffer's description of F. cana and is a better candidate for the species Fattig misidentified.

Figures

FIGURES 13–16. Dorsal habitus of Fidia species. 13. F.humeralis. 14. F. longipes. 15. F. marraverpa. 16. F. papillata.

FIGURES 42–47. Right prothoracic tibia (scale bar = 0.2 mm). 42. F. viticida ♂. 43. F. viticida ♀. 44. F. cana ♂. 45. F. convexicollis ♂. 46. F. longipes ♂. 47. F. rileyorum ♂.

FIGURES 93–95. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 93. F. longipes. 94. F. rileyorum. 95. F. delilahae.