Fidia Baly, 1863

Strother, M. S. & Staines, C. L., 2008, A revision of the New World genus Fidia Baly 1863 (Coleoptera: Chrysomelidae: Eumolpinae: Adoxini), Zootaxa 1798 (1), pp. 1-100 : 15-21

Key to the species of Fidia Baly

The following key, based on adult characters, separates all known species of Fidia . The first character given in each couplet is usually the character which is most easily visualized on undissected, pinned specimens. In order to properly view characters of the last abdominal sternum and pygidium of females, the abdomen must be dissected and placed in alcohol or glycerin. Supplemental characters and geographical distributions are given in most couplets to aid identification. Males and females are keyed separately to expedite the identification process and to accommodate single specimen and unisex series identifications. Females of most species can be keyed without difficulty; however, males are preferable because they often possess unique secondary sexual characters. Identifications based on female specimens should be confirmed with males, preferably by comparison of the genitalia with the figures provided.

1. MALE: disco-setae present on ventral surface of pro- and mesobasitarsi ( Fig. 42 ), often present on metabasitarsus as well in many Mexican and Central American species; individual setae usually not visible at dissection microscope magnifications (i.e., 25–50X) but indicated by "beaded" or hexagonal pattern or by faint multicolored sheen when brightly illuminated.......................................................... 2

1'. FEMALE: disco-setae never present on ventral surface of any basitarsus ( Fig. 43 ), ventral surface of basitarsus without "beaded" or hexagonal pattern and not imparting multicolored sheen when brightly illuminated ............................................................................................................................................ 23

2(1). Eyes not distinctly convex nor laterally protruding ( Fig. 55 ); northern Florida and adjacent Alabama and Georgia ............................................................................................... Fidia pedinops Strother n. sp.

2'. Eyes distinctly convex and laterally protruding ( Fig. 54 )....................................................................... 3

3(2). Pro- and mesotibial spurs robust, surface minutely rugulose (visible only at magnifications>25X) ( Figs. 42 and 45–47 ); species occurring predominantly north of Mexico (except F. clematis Schaeffer in extreme southern Texas and Mexico east of the Sierra Madre Oriental and F. marraverpa Strother n. sp. in Oaxaca, Mexico) ........................................................................................................................... 4

3'. All tibial spurs small ( Fig. 44 ), or rarely long and slender but always lacking visible surface sculpture at normal dissecting scope magnifications (i.e., 25–50X); species occurring predominantly south of the United States (except F. cana in central and southwestern Texas and F. humeralis in southeastern Arizona and southwestern New Mexico)................................................................................................... 12

4(3) Disco-setae on all basitarsi; mesepisternum glabrous; penis with broad, spatulate apical process ( Fig. 86 ) ......................................................................................................... Fidia marraverpa Strother n. sp.

4'. Disco-setae on pro- and mesobasitarsi only; mesepisternum distinctly pubescent; apex of penis with apical emargination ( Figs. 92–100 ) ........................................................................................................ 5

5(4') Pronotal punctation fine; apical emargination of penis without small medial tooth ( Figs. 92 and 96– 100 ) ......................................................................................................................................................... 6

5'. Pronotal punctation coarse; apical emargination of penis usually with small medial tooth ( Figs. 93–95 ) .............................................................................................................................................................. 10

6(5). Protibial spurs separated by much greater than basal width of one spur ( Fig. 45 ); dorsum of pronotum usually distinctly arched in lateral view ( Fig. 31 ); entirely black to dark red-brown with fine, silverywhite pubescence; apical emargination of penis rarely with feebly developed medial tooth; apical lobes slightly tapered to acutely rounded apex ( Fig. 92 ); southern Oklahoma and northern and central portions of east Texas ............................................................................... Fidia convexicollis Strother n. sp.

6'. Bases of protibial spurs contiguous or separated by less than basal width of one spur ( Fig. 42 ); dorsum of pronotum gently convex to nearly straight in lateral view ( Fig. 25 ); other characters variable......... 7

7(6'). Pro- and mesobasitarsi short, apical width subequal to length ( Fig. 53 ); occurring from extreme south Texas to Veracruz, Mexico, east of the Sierra Madre Oriental; apical emargination of penis deeply Ushaped, apical lobes long, feebly tapered to rounded apices ( Fig. 100 ) ............ Fidia clematis Schaeffer

7'. Pro- and mesobasitarsi long, length distinctly greater than apical width ( Figs. 54–55 ); widely distributed throughout eastern U.S.; apex of penis variously modified but not as above................................. 8

8(7'). Pro- and mesobasitarsi narrowly elongate, gradually widened from base to apex ( Fig. 53 ); apical emargination of penis semicircular, approximately as deep as wide ( Fig. 99 ) ................. Fidia viticida Walsh

8'. Pro- and mesobasitarsi broadly elongate, subequal in width for most of length ( Fig. 54 ); apical emargination of penis transverse, width much greater than depth ( Figs. 96–97 )........................................... 9

9(8'). Apical emargination of penis basally convex, apical lobes with acute apices ( Fig. 96 ); central Texas and New Mexico ................................................................................................... Fidia texana Schaeffer

9'. Apical emargination of penis basally concave or truncate, apical lobes with truncate to subtruncate apices ( Fig. 97 ); occurring in central U.S., excluding Texas ..................................... Fidia confusa Strother

10(5') All femora uniformly black to dark red-brown; beetle entirely black to dark red-brown; apical lobes of penis long, narrowly tapered ( Fig. 95 ); northwestern Alabama and extreme northeastern Mississippi ... .................................................................................................................. Fidia delilahae Strother n. sp.

10'. One or more pairs of femora bicolored, with base distinctly lighter than distal portion, or all femora less commonly entirely pale; beetle usually with light red-brown to flavous areas on head, elytra, and tibiae; apical lobes of penis usually not distinctly tapered; occurring throughout eastern half of U.S................................................................................................................................................................ 11

11(10').Protibial spurs separated by greater than length of one spur ( Fig. 46 ); width of apical emargination of penis approximately twice the width of one apical lobe ( Fig. 93 ); occurring predominantly east of the Appalachian Mountains .............................................................................. Fidia longipes (Melsheimer)

11'. Protibial spurs separated by slightly more than basal width of one spur ( Fig. 47 ); width of apical emargination of penis approximately four times width of one apical lobe ( Fig. 94 ); occurring west of the Appalachian Mountains ........................................................................... Fidia rileyorum Strother n. sp.

12(3'). Last abdominal sternum with shallow fovea, posterolaterally bordered by dense, brush of long, erect setae ( Fig. 59 ); other abdominal sterna lacking setal or structural modifications; mesepisternum pubescent; penis gently tapered to acutely rounded apex ( Fig. 88 ); Chiapas, Mexico and Suchitepequez, Guatemala................................................................................................. Fidia dichroma Strother n. sp.

12'. Abdominal sterna variable, but if last sternum has a fovea bordered by brush of setae, then medial area of second through fourth sterna with dense, transverse mat of short, stout setae ( Fig. 58 ); mesepisternum entirely glabrous or with several scattered setae, never as densely pubescent as mesepimeron .. 13

13(12').Medial area of second through fourth abdominal sterna with dense, transverse mat of short, stout setae and last sternum with broad, shallow, subcircular or transversely oval impression bordered laterally by dense brush of long, erect setae ( Fig. 58 ); apex of penis broadly convex, with small, "U" shaped notch ( Figs. 101 and 102 ) ............................................................................................................................... 14

13'. Abdominal sterna with or without various secondary sexual modifications but not fitting above description; apex of penis prolonged into an apical process of various sizes and shapes ( Figs. 81–91 ) or broadly emarginate with distinct apical lobes ( Fig. 80 ), but never convex with small apical notch... 15

14(13).Elytra coarsely, irregularly punctate-striate; beetle brassy-black with distinct gold, greenish, or purple metallic luster; elytral pubescence uniformly whitish to golden brown, setae suberect, ensate; legs fulvous, often with aeneous luster on femora; penis with apical notch approximately ¼ as long as distance between apex of penis and posterior margin of ostium with posterior angles prolonged into short, acute tooth-like projections, ( Fig. 101 ); Belize, El Salvador, and Guatemala....... Fidia guatemalensis Jacoby

14'. Elytral punctation confused, lacking striae; beetle glossy black; elytral pubescence composed of white, suberect, ensate setae on lateral aspect and sutural margins, and brownish or blackish, erect, hair-like setae on disc; penis with apical notch approximately ½ as long as distance between apex of penis and posterior margin of ostium, posterior angles not prolonged ( Fig. 102 ); known from a single specimen collected in central Costa Rica............................................................ Fidia costaricensis Strother n. sp.

15(13'). Medial area of third abdominal sternum with two small, round to subangulate, posteriorly directed lobes ( Fig. 60 ); apex of penis with long, spine-like apical process ( Fig. 85 ); Veracruz and Hidalgo, Mexico................................................................................................................. Fidia pedestris Lefèvre

15'. Second and third abdominal sterna not fitting above description; apex of penis variously modified .. 16

16(15').Penis with deep, semicircular apical emargination ( Fig. 80 ); known from a single specimen collected in Colima, Mexico................................................................................. Fidia comalensis Strother n. sp.

16'. Penis without apical emargination; apical process variously shaped ( Figs. 81–91 )............................ 17

17(16').Species small (2.86–3.32 mm.); brassy-black with distinctly contrasting fulvous tibiae; apex of penis with small, acutely angulate to sub-truncate apical process ( Fig. 83 ); declivitous part of penis distally widened in lateral view, widest at apex; known from Sierra Madre del Sur in central southern Mexico................................................................................................................................. Fidia tibialis Jacoby

17'. Not fitting above description, generally larger than 3.5 mm., but if smaller, then color of tibiae not distinctly different from that of dorsum; declivitous part of penis tapered distally in lateral view, widest proximal to apex ................................................................................................................................... 18

18(17').Apex of penis abruptly tapered from rounded posterolateral angles to bifurcate, spatulate process ( Fig. 81 ); known from single specimen collected in Mexico..................... Fidia dicelloposthe Strother n. sp.

18'. Apex of penis not fitting above description.......................................................................................... 19

19(18').Apex of penis broadly convex with small, acutely angulate apical process ( Fig. 82 ); color extremely variable, but often dark with red-orange humeral spot or lateral stripe on each elytron; southeastern Arizona and southwestern New Mexico; widely distributed in Mexico........... Fidia humeralis Lefèvre

19'. Apex of penis either not broadly convex ( Fig. 87 ) or if so, then apical process not small and acute ( Figs. 89–91 ); color uniformly very dark red-brown to black or, less commonly, fulvous to red-brown but never with humeral spot or lateral stripe ........................................................................................ 20

20(19').Species small (approximately 3.60 mm.); fulvous to red-brown; elytra with two broad, shallow impressions posteriad of intrahumeral callus ( Fig. 24 ); elytral pubescence predominantly light-colored, but with roughly M-shaped patch of darker setae crossing apical third of elytra; sides of penis gradually tapered, more abruptly so distad of ostium, to acutely rounded apex ( Fig. 87 ); Jalisco, Michoacan, and Oaxaca, Mexico......................................................................... Fidia xanthonioides Strother n. sp .

20'. Species larger than 4.0 mm.; very dark red-brown to black; elytra not fitting above description; apex of penis broadly convex with rounded posterolateral angles and well-defined apical process ( Figs. 89–91 ) .............................................................................................................................................................. 21

21(20').Pubescence of elytra and often pronotum with longer, hair-like, sub-erect to erect, brown to black setae in addition to shorter, ensate, recumbent to adpressed, white setae; white setae often arranged into distinct longitudinal vittae on elytra ................................................................. Fidia albovittata Lefèvre

21'. Pubescence of pronotum and elytra uniform in size, shape, and color................................................. 22

22(21').Species known only from central and southwest Texas; pubescence of pronotum often forming obscure to distinct medial longitudinal stripe when viewed with the un-aided eye or at very low magnifications (i.e., 6–12X); intrahumeral callus of elytron obsolete or only weakly developed; apical process of penis broad ( Fig. 89 ) .................................................................................... Fidia cana Schaeffer

22'. Species known only from Sierra Madre del Sur in central southern Mexico; pronotal pubescence never forming a medial longitudinal stripe; intrahumeral callus of elytron prominent; apical process of penis narrow ( Fig. 84 ) ....................................................................................................... Fidia spuria Lefèvre

23(1') Eyes not convex nor distinctly laterally protruding ( Fig. 34 ); northern Florida and adjacent Alabama and Georgia ............................................................................................... Fidia pedinops Strother n. sp.

23'. Eyes convex and distinctly laterally protruding ( Fig. 33 )..................................................................... 24

24(23').Last abdominal sternum with small, shallow to deep medial fovea ( Fig. 66 ) ..................................... 25

24'. Last abdominal sternum evenly convex, without medial fovea............................................................ 26

25(24) Species occurring in central Texas and New Mexico; generally larger than 6.8 mm. (6.52–8.08 mm.); dark red-brown to nearly black; elytral pubescence never arranged in distinct longitudinal rows .......... .............................................................................................................................. Fidia texana Schaeffer

25'. Species occurring throughout central U.S. but not known from Texas; generally smaller than 6.5 mm. (5.44–6.84 mm.); pale orange-brown to deep red-brown; elytral pubescence often arranged in feeble to distinct longitudinal rows .................................................................................... Fidia confusa Strother

26(24')Species occurring north of Mexico....................................................................................................... 27

26'. Species occurring in Mexico and Central America .............................................................................. 33

27(26).Mesepisternum entirely glabrous; known from southeastern Arizona and southwestern New Mexico; color extremely variable, but usually dark with distinct red-orange humeral spot or lateral stripe of variable width on each elytron ........................................................................... Fidia humeralis Lefèvre 27'. Mesepisternum sparsely to densely pubescent; occurring throughout eastern half of U.S. ................. 28

28(27')Pronotal punctation coarse.................................................................................................................... 29

28'. Pronotal punctation fine ........................................................................................................................ 31

29(28).Mesepisternum sparsely pubescent; pubescence of pronotum often forming obscure to distinct medial longitudinal stripe when viewed with the un-aided eye or at very low magnifications (i.e., 6–12X); occurring in central and southwestern Texas west of the 97th meridian ................. Fidia cana Schaeffer

29'. Mesepisternum densely pubescent; pubescence of pronotal disc uniformly distributed, never forming medial longitudinal stripe; occurring throughout eastern half of U.S. but known from Texas only east of the 97th meridian.............................................................................................................................. 30

30(29').All femora and tibiae unicolorous, very dark red-brown to black; (following two characters in both choices of this couplet visible only with abdomen dissected and placed in alcohol or glycerin): apical margin of last abdominal sternum with prominent, jagged-tipped medial process ( Fig. 63 ); lateral margin of pygidium with small, rounded preapical protuberance ( Fig. 71 ); northwestern Alabama and extreme northeastern Mississippi .............................................................. Fidia delilahae Strother n. sp.

30'. One or more pairs of legs partially fulvous to flavous, often with tibia and base of femur paler, distinctly contrasting with dark femoral apex, occasionally specimens from the southern U.S. with legs entirely fulvous to flavous; apical margin of last abdominal sternum with medial process poorly developed or absent ( Fig. 61 ); lateral margin of pygidium without preapical protuberance ( Fig. 72 ) ............. .............................................................. Fidia longipes (Melsheimer) or Fidia rileyorum Strother n. sp. (These two species are largely allopatric and are roughly separated by the Appalachian Mountains (Map 3), with F. longipes occurring predominantly to the east and F. rileyorum occurring only to the west. In areas of overlap, males are necessary to confirm identifications.)

31(28')Dorsum of pronotum usually distinctly arched in lateral view ( Fig. 31 ); beetle entirely black to very dark red-brown with dense, fine, silvery-white pubescence; known only from extreme southern Oklahoma and northern and central portions of east Texas ........................ Fidia convexicollis Strother n. sp.

31'. Dorsum of pronotum gently convex or nearly straight in lateral view ( Fig. 25 ), never distinctly arched; species widely distributed..................................................................................................................... 32

32(31').All basitarsi subtriangular, length 1.5–2 times apical width ( Fig. 57 ); species generally smaller than 5.5 mm. (4.56–5.60 mm.); occurring in extreme southern Texas ...................... Fidia clematis Schaeffer

32'. All basitarsi elongate, length 2.5–4 times apical width ( Fig. 56 ); species generally larger than 5.8 mm. (5.64–6.96 mm.); occurring throughout eastern half of North America north of Mexico........................ .................................................................................................................................. Fidia viticida Walsh

33(26').Mesepisternum densely pubescent; metafemur extremely narrow at base (Fig. 39); (following two characters in both choices of this couplet visible only with abdomen dissected and placed in alcohol or glycerin): apical margin of last abdominal sternum with small medial notch ( Fig. 68 ), pygidium with small apicomedial process ( Fig. 75 ); known from Guatemala and Chiapas, Mexico.............................. .................................................................................................................. Fidia dichroma Strother n. sp.

33'. Mesepisternum entirely glabrous or with few setae, never as densely pubescent as mesepimeron; shape of metafemur variable; apical margin of last abdominal sternum and apex of pygidium without apicomedial process ...................................................................................................................................... 34

34(33')Each elytron with feeble to distinct, subcircular to oval impression posterolaterad of postcallosal impression ( Figs. 16 & 24 ); setae of impressions, especially postcallosal impression, radiating from center; elytral pubescence uniform in size and shape, predominantly light-colored, but with brownish or blackish setae in jagged, transverse band or roughly M-shaped patch crossing elytra at apical third. .............................................................................................................................................................. 35

34'. Elytral disc evenly convex, sometimes with shallow lunate postcallosal impression, but never with additional impression posterolaterad of postcallosal impression; pubescence in postcallosal impression oriented as remainder of elytral pubescence; elytral pubescence unicolorous or with darker setae distinctly longer and finer than lighter setae ............................................................................................. 36

35(34).Metafemur basally extremely narrow for approximately half of length (Fig. 41); species large (approximately 6.0 mm.) and elongate ( Fig. 16 ); (two following characters in both choices of this couplet visible only with abdomen dissected and placed in alcohol or glycerin): apical margin of last abdominal sternum with two small papillae ( Fig. 65 ); apex of pygidium truncate with distinct posterolateral angles ( Fig. 73 ); known from single female collected in Hidalgo, Mexico............................................. .................................................................................................................. Fidia papillata Strother n. sp.

35'. Metafemur not as above (Fig. 40); species smaller (approximately 4.0 mm.) and more ovate ( Fig. 24 ); apical margin of last abdominal sternum without papillae ( Fig. 70 ); apex of pygidium broadly arcuate without posterolateral angles ( Fig. 74 ); Jalisco, Michoacan, and Oaxaca, Mexico................................. .......................................................................................................... Fidia xanthonioides Strother n. sp .

36(34').Elytral pubescence distinctly dimorphic, composed of shorter, ensate, recumbent, light-colored setae and longer, hair-like, suberect to erect, dark setae ............................................ Fidia albovittata Lefèvre

36'. Elytral pubescence uniform in size, shape, and color........................................................................... 37

37(36).Species small (3.06–3.36 mm.); dorsum entirely brassy-black with distinctly contrasting fulvous to red-brown tibiae; known from Sierra Madre del Sur in central southern Mexico... Fidia tibialis Jacoby

37'. Species larger than 3.80 mm.; not fitting above description................................................................. 38

38(37').Pronotum and elytra densely clothed with adpressed setae, surface sculpture not readily visible; pronotum densely, finely punctate-reticulate; occurring east of Sierra Madre Oriental from extreme southern Texas to central Veracruz, Mexico...................................................... Fidia clematis Schaeffer

38'. Pronotum and elytra not densely clothed with adpressed setae, surface sculpture readily visible; pronotal punctation not as above.................................................................................................................... 39

39(38').Apex of metatibia acutely subangulate in lateral view, widest some distance from apex ( Fig. 49 ); elytra coarsely, irregularly punctate-striate; beetle brassy-black with distinct gold, greenish, or purple metallic luster; legs fulvous, often with aeneous luster on femora; known from Belize, El Salvador, and Guatemala..................................................................................................... Fidia guatemalensis Jacoby

39'. Apex of metatibia broadly rounded in lateral view, widest at or near apex ( Fig. 48 ); other characters variable.................................................................................................................................................. 40

40(39').Species larger than 6.0 mm. (approximately 6.85 mm.); form robust in dorsal view ( Fig. 3 ); pronotum noticeably wider than long; elytra coarsely punctate-reticulate; dorsum very dark red-brown to blackish with faint blue-green luster; basal swelling type spermatheca ( Fig. 108 ); known from single female collected in Guerrero, Mexico..................................................................... Fidia chapini Strother n. sp .

40'. Species 5.5 mm. or smaller; form elongate in dorsal view; pronotal width and length subequal; elytral surface sculpture variable; basal arm type spermatheca ( Fig. 103 )...................................................... 41

41(40')Intrahumeral callus prominent with shallow, lunate postcallosal impression; elytral setae long, sparsely distributed; pronotal punctation moderately dense............................................................................... 42

41' Intrahumeral callus weakly developed or obsolete, without postcallosal impression; elytral setae short, moderately densely distributed; pronotal punctation dense and coarse................................................ 43

42(41) Apicomedial process of last abdominal sternum acute, with pointed apex ( Fig. 62 ); sides of pronotum distinctly arcuate; known from Sierra Madre del Sur in Guerrero and Oaxaca, Mexico......................... ................................................................................................................................. Fidia spuria Lefèvre

42'. Apicomedial process of last abdominal sternum subtruncate to feebly rounded at apex ( Fig. 64 ); sides of pronotum feebly arcuate; known from Sierra Madre Oriental in Hidalgo and Veracruz, Mexico....... ............................................................................................................................. Fidia pedestris Lefèvre

43(41')Humeral callus of elytron glabrous, darker than surrounding humeral area with surface polished; apicomedial process of last abdominal sternum apically rounded ( Fig. 67 ); known only from one ♂ and one ♀ collected in Oaxaca, Mexico............................................................. Fidia marraverpa Strother n. sp .

43'. Humeral callus of elytron setose or punctulate if setae abraded, same color as surrounding humeral area; apicomedial process of last abdominal sternum apically pointed ( Fig. 69 ); occurring along Sierra Madre Occidental in western Mexico from mountains of southeastern Arizona and southwestern New Mexico south to Oaxaca .................................................................................... Fidia humeralis Lefèvre

Figures

FIGURES 42–47. Right prothoracic tibia (scale bar = 0.2 mm). 42. F. viticida ♂. 43. F. viticida ♀. 44. F. cana ♂. 45. F. convexicollis ♂. 46. F. longipes ♂. 47. F. rileyorum ♂.

FIGURES 53–57. Pro- and mesobasitarsus (scale bar = 0.2 mm). 53. F. clematis ♂. 54. F. viticida ♂. 55. F. confusa ♂. 56. F. viticida ♀. 57. F. clematis ♀.

FIGURES 84–86. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 84. F. spuria. 85. F. pedestris. 86. F. marraverpa.

FIGURES 90–92. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 90. F. albovittata, Hidalgo, Mexico. 91. F. albovittata, Alta Verapaz, Guatemala. 92. F. convexicollis.

FIGURES 93–95. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 93. F. longipes. 94. F. rileyorum. 95. F. delilahae.

FIGURES 96–100. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 96. F. texana. 97. F.confusa. 98. F. pedinops. 99. F. viticida. 100. F. clematis.

FIGURES 31–34. 31. F. convexicollis lateral view of head and prothorax. 32. Graphops curtipennis schwarzi head. 33. F. rileyorum head. 34. F. pedinops head. Scale bar = 1.0 mm.

FIGURES 25–30. 25. F. rileyorum lateral view of head and prothorax. 26. F. convexicollis ventral view of prosternum. 27. Bromius obscurus lateral view of head and prothorax. 28. B. obscurus ventral view of prosternum. 29. F. convexicollis ventral view of mesosternum. 30. Aoria nigripes ventral view of mesosternum. Scale bar = 1.0 mm.

FIGURES 58–60. Male abdominal sterna, stippling indicates setose patches (scale bar = 1.0 mm). 58. F. guatemalensis. 59. F.dichroma. 60. F. pedestris.

FIGURES 87–89. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 87. F. xanthonioides. 88. F.dichroma. 89. F.cana.

FIGURES 101–102. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 101. F. guatemalensis. 102. F. costaricensis.

FIGURES 80–83. Distal part of male aedeagus, lateral and dorsal views (scale bar = 0.5 mm). 80. F. comalensis. 81. F. dicelloposthe. 82. F.humeralis. 83. F. tibialis. (abbreviations: ae- apical emargination; al- apical lobe; ao- apical orifice).

FIGURES 21–24. Dorsal habitus of Fidia species. 21. F. texana. 22. F. tibalis. 23. F. viticida. 24. F. xanthonioides.

FIGURES 61–70. Last visible abdominal sternum of female (scale bar = 1.0 mm). 61. F. rileyorum. 62. F.spuria. 63. F. delilahae. 64. F. pedestris. 65. F. papillata. 66. F. confusa. 67. F. marraverpa. 68. F. dichroma. 69. F. humeralis. 70. F. xanthonioides.

FIGURES 71–75. Female pygidium (scale bar = 1.0 mm). 71. F. delilahae. 72. F. rileyorum. 73. F. papillata. 74. F.xanthonioides. 75. F.dichroma.

FIGURES 13–16. Dorsal habitus of Fidia species. 13. F.humeralis. 14. F. longipes. 15. F. marraverpa. 16. F. papillata.

FIGURES 48–49. Leg characters (scale bar = 1.0 mm). 48. F. viticida left meso- and metathoracic tibiae, lateral view. 49. F. guatemalensis left metathoracic tibia, lateral view.

FIGURES 1–4. Dorsal habitus of Fidia species. 1. F. albovittata. 2. F.cana. 3. F. chapini. 4. F. clematis.

FIGURES 103–108. Female spermatheca, lateral view (scale bar = 0.5 mm). 103. F. convexicollis. 104. F. papillata. 105. F. xanthonioides, Jalisco, Mexico. 106. F. xanthonioides, Oaxaca, Mexico. 107. F. guatemalensis. 108. F. chapini. FIGURES 109–110. Male sperm guide, dorsal and lateral views (scale bar = 0.25 mm). 109. F. viticida. 110. F. cana.