Tyrannosaurus imperator, Paul & Persons & Van Raalte, 2022

Paul, Gregory S., Iv, W. Scott Persons & Van Raalte, Jay, 2022, The Tyrant Lizard King, Queen and Emperor: Multiple Lines of Morphological and Stratigraphic Evidence Support Subtle Evolution and Probable Speciation Within the North American Genus Tyrannosaurus, Evolutionary Biology 49 (2), pp. 156-179 : 19

publication ID	https://doi.org/ 10.1007/s11692-022-09561-5
DOI	https://doi.org/10.5281/zenodo.6324155
persistent identifier	https://treatment.plazi.org/id/039887A7-FFB1-DE57-FCB3-FD82FB97C376
treatment provided by	Tatiana
scientific name	Tyrannosaurus imperator
status	sp. nov.

Treatment

Tyrannosaurus imperator sp. nov. (Morphotype 1)

Etymology: Tyrant lizard emperor

Holotype: FMNH PR2081

Referred specimens: BHI 4182 , BHI 6231 , 6248 , HMN MB.R.91216 , MOR 1125 , MOR 1128 , RGM 792.000 , SDSM 12047 , TCM2001.90.1 , TMT v2222 , NMNNH P-3698?

Age and Stratigraphy: Late Maastrichtian, lower, lower middle and possibly middle Hell Creek and Lance, Laramie, Arapahoe, McRae? North Horn? Javelina?

Geographic distribution: Montana, Dakotas, Wyoming, New Mexico? Texas? Utah?

Diagnosis: Generally robust with an adult femur-length/ circumference ratio of 2.4 or less; usually two slender anterior incisiform dentary teeth.