Hemerodromia amazonensis , Câmara, J. T., Plant, A. R. & Rafael, J. A., 2014

Câmara, J. T., Plant, A. R. & Rafael, J. A., 2014, Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery I: new generic record and new species from Brazilian Amazon Basin, Zootaxa 3893 (2), pp. 209-231: 210-212

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Hemerodromia amazonensis

sp. nov.

Hemerodromia amazonensis  sp. nov.

( Figs 1–8View FIGURES 1 – 8)

Diagnosis. F 1 with basiventral denticle, with adjacent basal spines stronger ( Fig. 2View FIGURES 1 – 8); epandrium fused with hypandrium ( Fig. 8View FIGURES 1 – 8); surstylus complex, subovate lobe with spine-like setae at tip on inner face ( Fig. 6View FIGURES 1 – 8).

Description. Male ( Fig. 1View FIGURES 1 – 8). Head. Dark brown to black, antenna and mouthparts yellow with all setae whitish; ocellar triangle with 1 pair of proclinate setae; anterior ocellus larger. Eyes iridescent black, very narrowly separated on face bearing fine reclinate setulae. Frons with 3 setulae; small vertical setae contiguous with uniseriate postocular setae. Occiput bearing scattered fine hairs. Clypeus with rather dense short downwardly directed pile. Proboscis slightly curved, with setae yellow. Antenna with scape and pedicel bearing distinct short dorsal setulae; postpedicel about 1.5 X as long as wide, stylus ~ 0.8 X as long as postpedicel. Thorax ( Fig. 1View FIGURES 1 – 8). Elongate, slightly arched ventrally, dorsum yellow except brownish on prothoracic collar, narrowly behind postpronotal lobe, suture between katepisternum and meron; antepronotum with anterior margin almost straight; scutum with yellow setae, very small and fine except one notopleural. Legs ( Fig. 1View FIGURES 1 – 8). Whitish yellow. C 1 ~ 1.4 X longer than distance between C 1 and C2, 6X as long as wide with some pale dorsoapical setae. F 1 ~ 1.1 X as long as C 1, 3.5 –4.0X as long as wide; femoral formula 7 / 23 / 1 + 21 / 5, basal spines stronger ( Fig. 2View FIGURES 1 – 8); denticles confined to distal 0.7, 2 rows linear, black, without distal discontinuity, converging distally but divergent at tip. T 1 ~ 0.8 X as long as F 1, evenly curved, ventral face shallowly concave; with 2 rows of 17–21 sharply pointed spinose setae ventrally; dorsal ciliation of decumbent short pale setulae, denser distally; with small apicoventral extension on T 1 bearing strong apical spine.

Mid and hind legs slender with fine setae but T 3 with ‘comb’ of short setae posteroapically. Wing. Membrane faintly yellow, veins brownish; R 2 + 3 long and straight, not strongly curved towards C apically, joining C ~ 0.8 X distance between end of R 1 and R 4; R 4 + 5 fork angle acute (~ 60 °), distal to position of M 1 + 2 fork by ~ 2 X as long as R 4; R 5 and M 1 convergent distally; vein R 5 ~ 2 X as long as R 4; cell bm+dm short, ending at level of R 1, ~2.0X as long as cell br. Halter whitish yellow. Abdomen ( Figs 1, 3View FIGURES 1 – 8). Tergites 1–6 and 8 brownish; tergite 7 and sternites yellowish. Tergites with distinct setae on posterior margin; tergite 8 slightly shorter than sternite 8. Terminalia. Brown. Cercus with subbasal pointed projection, rounded at apex and distinctly setose on outer face ( Figs 4View FIGURES 1 – 8); right and left cerci close together dorsally ( Fig. 5View FIGURES 1 – 8). Epandrium wide, subtriangular ( Fig. 4View FIGURES 1 – 8), scattered setae on outer face, fused with hypandrium ( Fig. 8View FIGURES 1 – 8). Surstylus present, complex, subovate lobe, with spine-like setae at tip on inner face ( Figs 4, 6, 8View FIGURES 1 – 8). Hypandrium rather subtriangular ( Fig. 8View FIGURES 1 – 8), without distinct setae; gonocoxal apodeme projecting as narrow process just beyond anterior margin of hypandrium ( Fig. 4View FIGURES 1 – 8). Subepandrial sclerite wide, with subquadrate anterior projection and two subtriangular posterior processes ( Fig. 6View FIGURES 1 – 8). Phallus ( Fig. 7View FIGURES 1 – 8) sclerotized basally, extending just beyond hypandrium, with pointed dorsal projection apically and apical short membranous area with scattered spicules. Ejaculatory apodeme absent. Body length: 3.5 mm; wing length: 2.9 mm. Female. Unknown.

Type material. HOLOTYPE ♂ “ BRASIL, AM[azonas], S.[ão] Gabriel da Cachoeira, Ig.[arapé] Maturacá  , Malaise. 4.iv. 1995, Fernanda” ( INPA). PARATYPE: Idem, Manaus, PDBFF /WWF, Proj: Bert Klein, v. 1986, Malaise, Reserva 1301 (1 ♂, INPA).

Holotype condition. Right postpedicel and notopleural setae lost. Wing mounted on microslide. Terminalia in microtube with glycerin.

Etymology. The specific epithet refers to the State “Amazonas” where the specimens were collected.

Variation. Paratype with body length: 3.2 mm; wing length: 2.6 mm.

Remarks. Hemerodromia amazonensis  sp. nov. differs from other species principally by the fusion of the epandrium to the hypandrium.


Instituto Nacional de Pesquisas da Amazonia