Mortoniella umbonata, Blahnik & Holzenthal, 2008

Blahnik, Roger J. & Holzenthal, Ralph W., 2008, Revision of the Mexican and Central American species of Mortoniella (Trichoptera: Glossosomatidae: Protoptilinae), Zootaxa 1711 (1), pp. 1-72 : 64

Mortoniella umbonata , new species

Fig. 34

As previously discussed, Mortoniella umbonata is similar to a group of species, including M. carinula , M. opinionis , M. papillata , M. redunca , and M. sicula , all distinguished by having the inferior appendage invaginated mesally and with protruding, often apically acute, apicolateral projections, and also a phallicata with well developed dorsolateral processes. The apicolateral projections of the inferior appendage in M. umbonata are not as elongate as in the other species of this group, but more so than in either M. propinqua or M. florica which are otherwise similar in structure to members of this group. Among all of these species, M. umbonata is distinctively characterized by the paired, rounded, sclerotized processes extending from the lateral margin of the phallicata to the basoventral margin of the endophallic membrane.

Adult. Length of forewing: male 3.8–4.0 mm. Forewing with forks I, II, and III, hind wing with forks II and III. Overall color (in alcohol) light brown. Wing bar at anastamosis not evident, crossveins pale. Ventral process of abdominal segment VI (male) short, ventrally oriented, rounded or subtruncate apically, slightly constricted basally at anterior margin.

Male genitalia. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin forming rounded projection in dorsal half, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, dorsal excision narrow, much less than half width of segment. Tergum X with short, subacute mesal projection and projecting lateral lobes, lateral lobes short, subacute apically, slightly mesally curved. Inferior appendage without apicomesal projection, apicolateral projections short, stout, slightly dorsally curved apically, apices acute; appendage basolaterally, with margins undeveloped, lacking prominent projections on each side; mesal pockets of inferior appendage with apical processes short, dorsally curved. Paramere appendages very elongate, narrow, nearly uniform in width, slightly enlarged and scabrous apically, emerging from membranous lateral lobes, apices acute and slightly ventrally deflected. Dorsal phallic spine weakly sinuous in lateral view, apex distinctly upturned; as viewed dorsally, with apical half narrowed, apex rounded. Phallicata short, tubular, basal part relatively wide, dorsally with prominent, paired, rounded, dorsally protuberant projections on either side, laterally with sclerotized ridges, extending apicoventrally to form paired, rounded, umbonate processes. Endophallic membrane moderately elongate, with broadly rounded, membranous lobes dorsally, apicoventrally with single prominent, slender, curved spine.

Holotype male: PANAMA: Chiriquí: Guadalupe Arriba , 08°52'26"N, 082°33'13"W, 12–18.ix.1984, H. Wolda ( UMSP000118656 ) (alcohol) ( NMNH).

Paratypes: PANAMA: Chiriquí: same locality as holotype, 12–18.ix.1984, H. Wolda — 1 male (alcohol) ( NMNH) ; same locality, 6–12.xi.1985, H. Wolda — 1 male (alcohol) ( NMNH) .

Etymology. This species is named M. umbonata , from the Latin word umbo, a rounded protuberance, in reference to the paired ventral protuberances on the apicoventral margin of the phallicata, the most distinctive and diagnostic feature of this new species.

Figures

FIGURE 34. Mortoniella umbonata, new species. Male genitalia: A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal.