Ectrepesthoneura swolenskyi, BLAGODEROV & GRIMALDI, 2004

BLAGODEROV, VLADIMIR & GRIMALDI, DAVID, 2004, Fossil Sciaroidea (Diptera) in Cretaceous Ambers, Exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae, American Museum Novitates 3433 (1), pp. 1-76 : 39-41

Ectrepesthoneura swolenskyi , new species

Figures 50, 51 , Plate 5E

DIAGNOSIS: Sc long, meets R just before base of M3. Small radial cell relatively long. Gonocoxites with triangular ventral appendages bearing 2 long, dark apical spurs.

DESCRIPTION: Body length = 2.71 mm; wing length = 2.46 mm. Head: Ocelli three, from lateral ocellus to eye margin twice ocellus diameter. Flagellomeres cylindrical, length 1.5Χ width. Three palpomeres seen, apical one slender, length ratio 1:2:3.5. Thorax: Antepronotum with 2 setae, proepisternum with 4. Scutum with setae curved forward, lateral setae very long. Wing: All longitudinal veins except Sc with short setae. Costa ends beyond tip of R 5, one­half distance between tips of R 5 and M 1. Sc meets R just before M3 base. Length of R 1 2.2Χ that of r­m. Length of M3 2.7Χ that of r­m. Ratios of lengths of sections RS1, RS2, and RS3 1:2.3:7. RS1 oblique, 0.58Χ length of r­m. M3, M 1, and M 2 weak, M 1 slightly curved forward. M 3+4 and CuA curved gently caudad. Legs: Fore and mid tibiae longer than femora. Mid tibia with anterolateral, posterolateral, and posteromedial rows of short bristles, hind tibia with two rows of posterior setae. Basal third of mid tibia with large, dark, narrow sensory pit. Tibial spurs 1.3–1.7Χ tibial diameter. Abdomen. Ninth tergite large, conceals gonocoxites and gonostyli almost completely; caudal edge straight, without medial cleft. Ninth tergite straight caudally, without cleft, with numerous curved setae apically. Gonocoxites with triangular ventral appendages bearing 2 long, dark apical spurs. Gonostyli short, pointed, curved S­like.

MATERIAL: Holotype AMNH NJ 824 , male. USA: New Jersey, Sayreville; coll. S. Swolensky, 1997–1998 (mentioned in Grimaldi, 2000: fig. 48h).

ETYMOLOGY: The species epithet is patronymic for the late Steve Swolensky, who collected the specimen.

COMMENTS: The new species differs from other species of the genus in the structure of male genitalia, especially by ventral appendages of gonocoxites bearing dark sclerotized spurs.

Figures

Figs. 48–51. Ectrepesthoneura End. 48. E. succinimontana, n.sp., holotype PIN 3311/662. 49. Female genitalia. 50. E. swolenskyi, n.sp., holotype AMNH NJ­824. 51. Male genitalia.