Papunega magnifacies Gnezdilov et Le Cesne

Gnezdilov, V. M., Cesne, M. Le, Soulier-Perkins, A. & Bourgoin, T., 2015, New Guinean Issidae: description of new taxa in a poorly known island fauna (Hemiptera, Fulgoroidea), Zootaxa 3904 (1) : -

Papunega magnifacies Gnezdilov et Le Cesne , sp. nov.

( Figs 1–12)

Type material. ♂, holotype, Papua New Guinea, Wilhelm Mt, 1200 m, 5°44'28''S 145°19'38''E, 25.X.2012, Malaise trap. MNHN (EH) 22060.

Description. Metope wide, particularly above the clypeus, with thick subapical transverse carina and trace of median carina ( Figs 2, 12 ). Metope without sublateral carinae, but with two incomplete sublateral rows of tubercles. Lateral margins (keels) of metope raised, slightly leaf-shaped above the pedicel. Metopoclypeal suture 1 2

0,5 mm

0,5 mm 0,5 mm 3

1 mm 4 0.5 mm

6

5 0.5 mm 7

0.5 mm 8 9

0.5 mm 0.5 mm distinct, complete, widely convex. Postclypeus large. No ocelli. pedicel elongately cylindrical. Coryphe transverse, anterior margin almost straight, posterior margin obtusely angulately concave ( Figs 1, 10 ). Coryphe with two large tubercles near its posterior margin besides of median line. Lateral and posterior margins of coryphe keel-shaped. Rostrum reaching hind coxae. Pronotum longer than mesonotum, with trace of median carina, anterior margin keelshaped obtusely angulate, posterior margin almost straight. Paradiscal fields of pronotum very narrow behind the eyes. Paranotal lobes of pronotum rather narrow. Mesonotum with weak median and lateral carinae. Tegulae large. Fore wings elongate, slightly narrowing to rounded apices, without hypocostal plate. Basal cell narrowly oval. Radius with 2 branches (furcates near the basal cell), median with 5 main branches (first furcation near wing middle), cubitus anterior with 2 branches (furcates near wing middle) ( Fig. 3). Fore wing with many tranverse veins in its distal part. Clavus closed (postcubitus + first anal vein joining cubitus posterior before apex of clavus). Hind wings 3-lobed ( Fig. 4), with coupling lobe. Posterior branch of cubitus anterior (CuA3) and CuP terminating in deep clef of wing margin and becoming flattened apically ( Fig. 4). Anterior branch of first anal vein (A1 1) joining Pcu in middle. Posterior branch of first anal vein (A1 2) running to weak cleft of wing margin. Hind tibia with 2 lateral spines distally ( Fig. 12 ) and 7 spines apically. First metatarsomere with 2 lateral and 5 intermediate (in entire row) spines apically.

Coloration. General coloration light brown yellowish ( Figs 10–12 ). Metope in frontal view with 4 peculiar bands (from its upper margin to clypeus): first band dark brown with a transverse paler brown band in middle (upper one, between the eyes); second thinner whitish band (between ventral sides of the eyes); from dark brown to black widest third band between antennae; whitish thinner straight fourth band above the metopoclypeal suture ( Fig. 12 ). Postclypeus dark brown to black frontally, with light brown yellowish lateral parts. Anteclypeus totally black. Apices of rostrum and spines of legs black. Fore wings distally with dark brown cells ( Figs 10, 11 ). Hind wings with dark brown veins. Hind coxae with large black spots. Fore and middle trochanters dark brown yellowish. Hind trochanters dark brown to black. Hind femorae with inner surfaces dark brown. Fore and middle legs with third tarsomeres dark brown. Claws of fore and middle legs dark brown. Claws of hind legs brown.

Male genitalia ( Figs 5–9). Anal tube elongate, narrow (2.5 times as long as wide medially) in dorsal view ( Fig. 7). Anal column short ( Fig. 5). Pygofer with slightly concave hind margin in lateral view ( Fig. 5). Each dorsolateral phallobase lobe with a simple short subapical hook-shaped process directed apically ( Fig. 8). Aedeagus with pair of furcated ventral hooks directed basally (one long branch almost reaching phallobase bottom and another one short) ( Fig. 8). Ventral phallobase lobe wide, enlarged subapically in ventral view ( Fig. 9). Gonostyles with straight hind margin; capitulum on long neck; lateral tooth small, apical tooth distinct ( Fig. 6).

Total length. Male—5.6 mm.

Etymology. Species name is derived from the combination of Latin "magna" (large) and "facies" (face) referring to the wide metope of the species.

Figures

FIGURES 10 – 12. Papunega magnifacies gen. et sp. nov., 10 — habitus, dorsal view; 11 — habitus, lateral view; 12 — frontal view.