Tephritis maccus

Namin, Saeed Mohamadzade & Korneyev, Severyn V., 2012, Tephritis azari, a New Fruit Fly (Diptera: Tephritidae) from Iran and Azerbaijan, with a Key to the Tephritis maccus Species Group, Zootaxa 3590, pp. 79-85 : 83-84

Key to the species of the Tephritis maccus species group

1 Wing pattern reticulate, radiate or banded; cell r1 with two or three hyaline spots or indentations (if with two spots in r1 and wing apex and cell m with narrow dark “rays”, then cell dm mostly dark with 3–5 hyaline spots); flagellomere 1 apically rounded; aculeus gradually tapered to the apex; with preapical steps or without them [spermathecae variabe]..... Other Tephritis species

- Wing pattern radiate (apex and cell m with narrow dark “rays”), cell r1 either without hyaline spots ( Figs. 14–15 ) or with only one large hyaline spot or indentation (if rarely with two spots, then cell dm mainly hyaline with at most two dark rays crossing it); flagellomere 1 apically acute or pointed; aculeus moderately broad, sharply tapered subapically to short narrow apex ( Figs. 4–5 ); spermathecae pear-shaped ( Fig. 8 ).................................................................... 2

2 Cell r1 without large hyaline spots; apical fork with base broadly connected to main pattern; cell dm dark with one hyaline spot ( Figs. 14–15 ); Iraq, Iran and Turkmenistan..................................................... T. urelliosomima

- Cell r1 with one large (and sometimes with one smaller) hyaline spot; apical fork with base narrowly connected to main pattern; cell dm hyaline with 2–3 transverse brown rays.............................................................. 3

3 Brown ray in cell dm basal to level of r-m long, reaching middle of cell cua1 ( Fig. 13 and Korneyev & Dirlbek 2001, Figs. 2–3 ). Female: Cell r4+5 at level of dm-cu without or with small hyaline spot; aculeus broad, about 2.5 times as long as wide ( Korneyev & Dirlbek 2001, Fig. 8 ). Male wing with cell r1 usually including 2 hyaline spots: large one touching R2+3 or entering cell r2+3; Spain, France, Italy, Iran, Kyrghyzstan and Afghanistan.................................. T. maccus

- Short brown ray in cell dm basal to level of r-m absent or at most reaching vein Cu1, usually reaching only mid-width of cell dm. Female: cell r4+5 at level of dm-cu with large hyaline spot; aculeus narrower, about 3 times as long as wide. Male wing with large hyaline spot in r1 cell always penetrating to at most mid-width of cell r2+3..................................... 4

4 Brown area around R2+3 apex with broad U-shaped darker spot (including small hyaline spot at R2+3 apex) darker than remaining pattern. Both male and female with 2 hyaline spots in cell r1 ( Gentilini et al., 2006: Figs. 30–33, 36–38). Fossil species from Upper Miocene of Italy................................................................ T. sophus

- Brown area around R2+3 apex as dark as remaining pattern. Female with one spot, male with 2 hyaline spots in cell r1 ( Figs. 1–2 , 12 ). Recent species. Azerbaijan and Iran............................................................... T. azari

Figures

FIGURES 13 – 15. Tephritis maccus (SMNC) (13) and T. urelliosomima (SIZK) (14 – 15): 13 – 14 Ƥ, total view, left 15 wing.

FIGURES 1 – 8. Tephritis azari sp. n., 1 wing pattern in male (paratype) 2 wing pattern in female (Holotype) 3 head in profile 4 aculeus 5 aculeus tip 6 epandrium 7 male terminalia 8 spermathecae.

FIGURES 9 – 12. Tephritis azari sp. n., paratype Ƥ (Azerbaijan, SIZK), 9, total view, dorsal 10 total view, right 11 abdomen 12 wing pattern.