Rana, Linnaeus, 1758

Grismer, L. Lee, Neang, Thy, Chav, Thou, Wood, Perry L., Jr & Oaks, Jamie R., 2008, Additional Amphibians And Reptiles From The Phnom Samkos Wildlife Sanctuary In Northwestern Cardamom Mountains, Cambodia, With Comments On Their Taxonomy And The Discovery Of Three New Species, Raffles Bulletin of Zoology 56 (1), pp. 161-175 : 164-167

Rana cf. milleti Smith, 1921

( Fig. 8 )

Material examined. – LSUHC 7828 , 7874 : approximately 5 km north of Camp 1 (12º09.140'N 103º07.719'E), 6 and 8 Aug.2006 , respectively; LSUHC 7892–97 : Camp 2, 9 Aug.2006 .

Remarks. – A series of eight males (SVL 45–48 mm) match the description of this species by Smith (1921) and more closely the expanded descriptions by Inger et al. (1999) and Chuaynkern et al. (2004) in having small round vocal sacs below the angle of the jaws with openings in the corners of the mouth; a strong dorsolateral fold; granular skin on back with many small tubercles capped by spinules with larger tubercles on the side; smooth skin on the belly; and a wide, dark stripe extending from the tip of the snout, through the eye, to the tympanum. The specimens differ in having more extensive webbing on the hind foot that extends to over onehalf the length of the toes and the SVLs of males are greater TABLE 1: Checklist of the herpetofauna of the northwestern Cardamoms. Literature sources for species previously reported from the northwestern Cardamoms are listed in Grismer et al. (2007a).

TABLE 1: Cont'd.

than that reported by Chuaynkern et al. (2004; 45–48 mm vs. 32.6–41.7 mm). Stuart & Emmett (2006) noted similar webbing to that reported here for two specimens from the central Cardamoms. This series differs from those of Inger et al. (1999), Smith (1921), and Stuart & Emmett (2006) in that all ventral surfaces are immaculate as opposed to having dark mottling and they differ from the description of Chuaynkern et al. (2004) in that the posterior portion of the thigh is lightly mottled as opposed to being a uniform brown.

All specimens were collected in swampy areas that remain wet throughout the year. They were located during the day and night by their calls. Chuaynkern et al. (2004) report this species from the northeastern Cardamoms.

Figures

Fig. 8. Rana cf. miletti from Camp 1.