Trichadenotecnum krucilense Endang, Thornton & New

Yoshizawa, Kazunori, Lienhard, Charles & Abd, Idris, 2014, Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae), Zootaxa 3835 (4), pp. 469-500 : 497-498

Trichadenotecnum krucilense Endang, Thornton & New

( Figs 1U , 29 )

Trichadenotecnum krucilense Endang, Thornton & New, 2002: 165 ; Endang & New, 2005: 23.

Specimen examined. [ Malaysia] 1 male, Gnung Berembun , Path 3, Cameron Highlands, Pahang, 14.vii.2003, HK .etal (canopy fogging, St. 6).

Remarks. The species was originally described from Java, Indonesia, later also recorded from Sumatra, Indonesia, and is here recorded from Peninsula Malaysia for the first time. The variation of the hypandrial structure, three (Javanese) or four (Sumatran) hypandrial spikes, has been pointed out by Endang & New (2005). The presently examined specimen has four hypandrial spikes. Judging from the original hypandrial illustration published by Endang et al. (2002: fig. 333), missing of some hypandrial parts on the left side of the specimen seems evident; on that side only one spike is figured whereas two on the other side. It is likely that the variation pointed out by Endang & New (2005) is actually an artifact caused by dissection or preparation.

The hypandrium of this species ( Fig. 29C ) is unique, very different from that of any other known species. Absence of the hypandrial median tongue may indicate its close affinity with T. apertum and its close relatives. In the original description of this species, the phallosome is illustrated in highly deformed condition, but the present examination clearly shows that this species has the pseudo-parameres on the phallosome ( Fig. 29D ). The presence of this structure is a synapomorphy of the sexpunctatum, medium and New World species groups (see Yoshizawa et al., 2008) so that assignment of T. krucilense into the majus group is excluded. However, this species cannot be assigned to the sexpunctatum, medium or New World species groups because of lack of apomorphies characterizing any of these species group so that a new species group for its own is proposed here. Females of this species are unknown to date.

Figures

FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male.

FIGURE 29. Male terminalia of Trichadenotecnum krucilense. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.