Trichadenotecnum endangae Yoshizawa & Lienhard, 2014

Yoshizawa, Kazunori, Lienhard, Charles & Abd, Idris, 2014, Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae), Zootaxa 3835 (4), pp. 469-500 : 481-483

Trichadenotecnum endangae Yoshizawa & Lienhard , sp. n.

( Figs 1I , 11 , 12 )

Holotype. Male, Malaysia, Gunung Brinchang, Cameron Highlands , Pahang, 11.iii.2003, K. Yoshizawa ( UKM).

Paratype. 1 female, Malaysia, Gunung Jasar, Cameron Highlands , Pahang, 9.iii.2003, KY ( UKM) .

Description. Male. Head. Yellowish white in ground color; vertical markings blackish brown, each marking fused with neighbors; with pair of blackish brown markings anterior to vertical markings; orbital markings blackish brown; coronal suture black; epicranial suture broadly bordered with blackish brown band dorsally; frons mostly blackish brown, with pair of dorsal white spots and pair of lateral white bands; eye black, IO/D= 1.3; ocelli white, ocellar field black; gena mostly blackish brown; postclypeus mostly blackish brown, dorsally with six white bands; anteclypeus blackish brown. Antenna brown, scape and pedicel blackish brown. Mouthparts brown.

Thorax. Prothorax blackish brown. Mesonotum blackish brown, medially with trifurcated white marking. Metanotum blackish brown, medially with white spot. Meso- and metapleuron brown except for white membranous regions.

Legs. Mostly blackish brown, distal half of ventral surface of fore femur and distal tip of hind femur white, tibiae paler.

Forewing ( Fig. 1I ). Extensively covered with tiny spots, dense basally from basal band, sparse distally from basal band. Distal spot in cell a1 distinct, basal spots obscure. Opposing spots in cell r obscure. Basal band narrow but distinct, broadly interrupted above M-Cu fork, narrowly interrupted above CuP vein. Distal band narrow but distinct. Spot on roof of cell m3 faint, only distinct at top of areola postica. Submarginal spots distinct in cells r3 to m2, obscure in other cells. Marginal clouds almost absent. Hindwing hyaline; veins brown.

Abdomen dark brown, with irregular white spots.

Terminalia. 8th sternum with single short triangular sclerite fused to hypandrium ( Fig. 11 AC). Clunium ( Fig. 11A ) without well developed clunial arm. Epiproct ( Fig. 11 AB) conical in shape, shorter than wide, extended posteriorly. Paraproct ( Fig. 11A ) with well developed complex basal process; distal lobe short and rounded; distal process straight, directed posteriorly. Hypandrium ( Fig. 11C ) asymmetrical, posteromedially with broad membranous region; long, narrow, weakly sinuous and distally pointed process arising from right side; posterolateral corner with saw-like posterior projection. Phallosome ( Fig. 11D ) with rugous posterior margin, medially with deep notch, laterally rounded, posteriorly with broad anterior process.

Measurements. B 2.4, Fw 3.5, Hw 2.6.

Female. General morphology almost as in male. IO/D = 2.5.

Genitalia. Egg guide ( Fig. 12A ) broad basally, strongly constricted to middle and slightly broadened to slightly arched distal margin, laterally with membranous regions; body of subgenital plate broad, anteriorly with broad membranous regions and anteromedially with deep V-shaped membranous region. Gonapophyses ( Fig. 12B ). Ventral valve long. Dorsal valve with long distal process. External valve lacking posterior lobe, narrow. Spermapore plate ( Fig. 12C ) with strongly pigmented area posteriorly.

Measurements. B 2.3, Fw 3.1, Hw 2.4.

Etymology. The species epithet honors the memory of Endang Sri Kentjonowati, author of several important contributions to the knowledge of Indonesian Psocoptera , especially of the family Psocidae . She died, much too young, on 29 April 2010.

Remarks. This species is very close to T. bromoense Endang, Thornton & New, 2002 from Java, Indonesia, but differs from it by the shape of the paraproctal lateral process ( Fig. 11A ). In the original description, the long hypandrial process of T. bromoense arises from the left side. However, in ventral (external) view, the process arises from the right side in T. endangae ( Fig. 11C ), as well as in close relatives such as T. godavarense New, 1971 and T. sclerotum New, 1978 . Therefore, it is very likely that the illustration of the hypandrium of T. bromoense is drawn in dorsal (internal) view or reproduced in reversed position.

Figures

FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male.

FIGURE 11. Male terminalia of Trichadenotecnum endangae. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.

FIGURE 12. Female genitalia of Trichadenotecnum endangae, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate.