Paratrizygia conformis Tonnoir 1929

Amorim, Dalton De Souza, Oliveira, Sarah Siqueira & Mcalister, Erica, 2011, The identity of Paratrizygia conformis Tonnoir (Diptera, Mycetophilidae), with comments on its systematic position, Zootaxa 2892, pp. 47-52 : 48-50

Paratrizygia conformis Tonnoir 1929

Paratrizygia conformis Tonnoir 1929: 605 , text–fig. 4 (male hypopygium), plate xxiii, fig. 15 (wing) [wrongly indicated as plate xxiii, fig. 14].

Redescription of the holotype ( Figs. 1–3 ).

Male. Head. Dark brown. Both antennae broken on the holotype, one of them with the first flagellomere; scape and first flagellomere dark brown, pedicel slightly lighter. Mid ocellus very small, present, lateral ocelli more than twice the mid ocellus, almost touching the eye margin. Eye slightly emarginated above the antenna. One of the maxillary palpi broken, the other dark brown, basal palpomere very small, apical ones increasingly longer, last palpomere almost twice length of penultimate. Thorax. Scutum dark brown, moderately arched, covered with scattered setae, no clear row of acrostichals or dorsocentrals, a row of slightly stronger supra–alars and some strong prescutellars. Scutellum dark brown, a pair of longer scutellar setae laterally, a pair of setae half of its length more mesally and a number of smaller setae. Pleural sclerites dark brown, except for the metepisternum, light brown. Pleural membrane brown. Pronotum and proepisternum densely setose, with some stronger setae. Anepisternum bare; katepisternum more or less squared ventrally. Mesepimeron reaching ventral margin of thorax, bare. Laterotergite slightly projected, with a group of 7–8 setae. Mediotergite slightly curved in profile, ventral half with a pair of patches of longer setae and some setulae. Haltere pedicel brown yellow, knob darker. Legs light brown, elongate. Mid and hind tibiae with dorsal and lateral regular rows of longer, black setae, mid tibia with about four setae, hind tibia about 9. Mid and hind tarsomeres with a row of shorter, black ventral setae, first tarsomere more than twice the length of second tarsomere; Tibial spurs 1:2:2, outer spur length almost three times width of tibial apex, inner spur about half the length of the outer one. Tarsi mostly missing in the holotype (both front and one of the mid tarsi entirely missing, distal articles missing in the other mid and both hind tarsi). Wing ( Fig. 1 ). Membrane translucent on basal two thirds, light brown apically, especially at anterior margin and along posterior margin of CuA; membrane dense and regularly covered with microtrichia, macrotrichia present, especially distally and posteriorly, basal cells with few or no macrotrichia. Sc complete, normally sclerotized, reaching C just beyond base of Rs, setose; sc–r absent. C ending before wing apex, extending about a fourth the distance between R5 and M1. First sector of Rs clearly oblique, short, devoid of setae, more than twice as long as r–m. R1 relatively long, reaching C at apical third of wing, with dorsal and ventral macrotrichia; R4 present, devoid of setae, sector of Rs between origin of r–m to base of R4 about three times length of r–m; R5 short, reaching C almost at level of apex of M4, with dorsal and ventral macrotrichia; r–m oblique, well sclerotized, devoid of setae. M1+2 unforked, with fine setae only dorsally; an unattached vein between M1+2 (possibly M4) and CuA, slightly curved, beginning before apex of R1, with fine setae only dorsally. CuA unforked, normally sclerotized, with fine setae only dorsally, except for the very base, in which setulae are absent. A1 no visible (even though referred to in original description). Abdomen. Abdomen dark brown, setose, slender. S8 slender to the apex, longer than wide, T8 short and wide. Terminalia ( Figs. 2– 3 ). Terminalia light brown. Gonocoxites setose, not too elongate, in lateral view not possible to verify the median suture, clearly no distal extension beyond level of insertion of gonostyles. Gonostyles elongated, more or less slen- der, with two very strong setae basally and about 8–10 slender spines apically; aedeagus not visible; gonocoxal apodeme not visible; T9 elongated, setose, with two pairs of regular rows of spines on an distal fold facing ventrally; cerci lobose, small.

Comments. Matile (1998, Fig. 5) illustrated the male terminalia of P. conformis and stated that he examined the “type-species” of Paratrizygia ( Matile 1998: 387) . His focus in that paper was the genus Afrocnemia Matile and he did not include a list of specimens examined from other genera. He probably had access to the holotype at the NHM, but did not state this clearly in his comments about the material examined. His illustration of the male terminalia of P. conformis is in ventral view ( Matile 1998, Fig. 5) and some details, especially of the tergite 9, were not included.

Figures

FIGURE 1. Paratrizygia conformis Tonnoir, holotype. Wing. Scale 1 mm.

FIGURE 2. Paratrizygia conformis Tonnoir, holotype. Male terminalia, lateral view. Scale 0.2 mm. Abbreviations: Ce, cercus; Gc, gonocoxite; Gs, gonostyle; S 8, sternite 8; T 8, tergite 8; T 9, tergite 9.

FIGURE 3. Paratrizygia conformis Tonnoir. Male terminalia, lateral view. Scale 0.2. Abbreviations: Ce, cercus; Gc, gonocoxite; Gs, gonostyle; T 8, tergite 8; T 9, tergite 9; tsp tergite 9 spines.