COCHYLINI Guenée, 1845

Byun, Bong-Kyu & Li, Cheng-De, 2006, Taxonomic review of the tribe Cochylini (Lepidoptera: Tortricidae) in Korea and northeast China, with descriptions of two new species, Journal of Natural History 40 (13 - 14), pp. 783-817 : 785-787

Tribe COCHYLINI Guenée, 1845

5Cochylidii Guenée 1845, p 297.

5 Cochylidae Morris 1868, p 209 .

5 Phaloniidae Meyrick 1895, p 543 .

5 Cochylini Kuznetsov and Stekolnikov 1973, p 21 .

Key to the adults of the genera of the tribe Cochylini

1. Apex of forewing rounded, medial fascia not oblique inwardly Phtheochroa Stephens – Apex of forewing narrowed, medial fascia oblique inwardly....... 2

2. Basal fascia in forewing broad, well developed.... Phalonidia Le Marchand

– Basal fascia in forewing weak or absent............. 3

3. Subterminal fascia distinct along termen............ 4

– Subterminal fascia very weak or absent............. 5

4. Median fascia of forewing expanded, vein R 5 running to costa or apex.......................... Cochylis Treitschke

– Median fascia of forewing subtriangular, vein R 5 running to termen........................... Eupoecilia Stephens

5. Median fascia curved inwardly near middle, several pretornal markings present. 6

– Median fascia nearly straight towards middle of dorsum........ 7

6. Distance between R 1 – R 2 in forewing shorter than R 2 –R 3 ... Aethes Billberg

– Distance between R 1 –R 2 in forewing similar or slightly longer than R 2 – R 3 ................ Gynnidomorpha Turner

7. Vein R 5 in forewing running to costa....... Cochylidia Obraztsov

– Vein R 5 in forewing ended at termen............. 8

8. Vein M 3 and vein Cu 1 of hindwing separate.... Cochylimorpha Razowski

– Vein M 3 and vein Cu 1 of hindwing stalked...... Eugnosta Hübner

Key to the genera of Cochylini based on male genitalia

1. Uncus reduced or absent; aedeagus usually with numerous cornuti, sometimes with one or two longer cornuti............... 2

– Uncus present, longer than socii; aedeagus with several cornuti Phtheochroa Stephens 2. Sacculus and valva not fused................ 3

– Sacculus and valval bases fused............... 4

3. Valva broad, elongated, complicatedly concaved outwardly; apex of aedeagus shortly protruded, curved inwardly......... Cochylis Treitschke

– Valva narrow, short, well sclerotized; apex of aedeagus slender Cochylidia Obraztsov

4. Socii broad, mostly triangular; ventral and costal margins of valva nearly parallel................ Eupoecilia Stephens

– Socii narrow, slender, hook-shaped; ventral and costal margins of valva not parallel 5

5. Medial process of transtilla broad, short, not bifurcate apically...... 6

– Medial process of transtilla thin, long, bifurcate apically........ 7

6. Valva orbicular; sacculus weakly developed; socii erect........ 8

– Valva broad basally, narrowed towards apex; sacculus well sclerotized; socii droop- like................ Cochylimorpha Razowski

7. Tegumen narrow, sacculus weakly sclerotized.... Gynnidomorpha Turner

– Tegumen broad, sacculus well sclerotized..... Phalonidia Le Marchand

8. Socii narrow curved.............. Aethes Billberg

– Socii thick, erect.............. Eugnosta Hübner

Key to the genera of Cochylini based on female genitalia

1. Antrum membranous, ostium bursae modified.......... 2

– Antrum well sclerotized, ostium bursae simple.......... 5

2. Appendix bursae of corpus bursae usually present; corpus bursae with ornamentation, often indistinguishable from ductus bursae....... 3

– Appendix bursae of corpus bursae not present, corpus bursae simple, distinguishable from dusctus bursae................. 4

3. Ductus bursae always wrinkled and complicately sclerotized near ostium bursae; ostium bursae relatively narrow with a broad and complicated antrum......................... Cochylidia Obraztsov

– Ductus bursae usually simple, sometimes wrinkled; ostium bursae very large with simple antrum............... Cochylis Treitschke

4. Ductus bursae strongly twisted before corpus bursae... Phtheochroa Stephens

– Ductus bursae not twisted before corpus bursae.......... 6

5. Corpus bursae semiovate, with numerous tiny spinules; ductus bursae long................ Gynnidomorpha Turner

– Corpus bursae long, sack-like, without spinules, indistinguishable from ductus bursae............... Phalonidia Le Marchand

6. Corpus bursae with a large sclerite signum............ 7

– Corpus bursae without a sclerite............... 8

7. Corpus bursae wrinkled near ductus bursae, ductus bursae broad, very short; accessory bursae usually arising from base of corpus bursae... Aethes Billberg

– Corpus bursae simple near ductus bursae, ductus bursae fairly long; accessory bursae usually arising from corpus bursae..... Cochylimorpha Razowski

8. Ductus bursae usually with numerous spines..... Eupoecilia Stephens

– Ductus bursae usually without spines........ Eugnosta Hübner