Thienemanniella

Fu, Yue, Saether, Ole & Wang, Xinhua, 2010, Thienemanniella Kieffer from East Asia, with a systematic review of the genus (Diptera: Chironomidae: Orthocladiinae), Zootaxa 2431, pp. 1-42 : 32-34

Key to known males of Thienemanniella

1. Eyes bare, antennal apex distinctly rounded................................................................................................................. 2

- Eyes pubescent or hairy, antennal apex notched or slightly rounded ........................................................................... 7

2. Inferior volsella absent ....................................................................................................... T. flavescens Edwards, 1929

- Inferior volsella present ................................................................................................................................................ 3

3 Antenna with 9–10 flagellomeres ................................................................................................................................ 4

- Antenna with 11–13 flagellomeres ............................................................................................................................... 5

4. Antenna with 9 flagellomeres, inferior volsella rounded .................................. T. togamijika Sasa et Okazawa, 1992

- Antenna with 10 flagellomeres, inferior volsella hook-shaped ................................................. T. okigrata Sasa, 1990

5. Inferior volsella rectangular and projecting................................................................................ T. partita Schlee, 1968

- Inferior volsella nearly triangular and along the inner margin of gonocoxite ............................................................. 6

6. Antenna with 12 or 13 flagellomeres, scutellum black in male ( Tokunaga, 1936) ...... T. nipponica ( Tokunaga, 1936)

- Antenna with 11 or 12 flagellomeres, scutellum yellow in male ( Tokunaga, 1936) T. flaviscutella ( Tokunaga, 1936)

7. Inferior volsella absent ................................................................................................................................................. 8

- Inferior volsella present ............................................................................................................................................. 10

8. Superior volsella hook-shaped; transverse sternapodeme without oral projection; phallapodeme straight and short .. ....................................................................................................................................................... T. hainanensis sp. n.

- Superior volsella triangular, transverse sternapodeme with oral projections; phallapodeme more or less curved and relatively long ............................................................................................................................................................... 9

9. Antenna with 11 flagellomeres, AR 0.72–0.90; third palpomere with digitiform lobe ......... T. similis ( Malloch) 1915

- Antenna with 12 flagellomeres, AR 0.25–0.28; third palpomere without digitiform lobe.................... T. absens sp. n.

10. Inferior volsella narrow, digitiform to spatulate, sometimes with basal apodeme; or broad and low ........................ 11

- Inferior volsella well developed, rectangular to triangular......................................................................................... 20

11. Superior volsella completely fused with the inner of gonocoxite ........................... T. medialis Sublette & Sasa, 1994

- Superior volsella present, well developed or partly fused with the inner of gonocoxite............................................ 12

12. Inferior volsella digitiform with an apodeme, bare, phallapodeme straight ............................................................... 13

– Inferior volsella not digitiform, most often carrying thin glandular setae, phallapodeme curved ............................. 14

13. Superior volsella with a flat rim along the gonocoxite narrowly and completely fused .. T. antennata Freeman, 1953

- Superior volsella mostly triangular and fused only anteriomedially ............ T. lobapodema Hestenes & Saether, 2000

14. Inferior volsella placed basally or medially on gonocoxite........................................................................................ 15

- Inferior volsella very low and placed caudally on gonocoxite ................................................................................... 18

15. AR 0.52–0.58, inferior volsella very broad and rounded ............................................................................................... ........................................................................................................................... T. akagiquarta Kikuchi & Sasa, 1994

- AR 0.20–0.30, inferior volsella digitiform or narrow................................................................................................. 16

16. Gonocoxite with large concavity for gonostylus, tergite IX with many long setae........................................................ ......................................................................................................................... T. cavata Fu, Hestenes & Saether, 2010

- Gonocoxite without such concavity for gonostylus, tergite IX without many long setae .......................................... 17

17. Antenna with 8 flagellomeres; tergite IX with a small, flexible, triangular hump-like extension; apex of phallapo- deme bent 90° caudally. ............................................................................................................ T. fuga Lehmann, 1979

- Antenna with 10 flagellomeres; tergite IX without hump-like extension; apex of phallapodeme slightly curved ........ .............................................................................................................................................. T. lineola Freeman, 1953

18. Antenna with 9 flagellomeres; gonostylus expanded; superior volsella more or less digitiform and projecting........... ...................................................................................................................................... T. nagaramaculata Sasa, 1989

- Antenna with 10 flagellomeres; gonostylus not expanded; superior volsella narrow and not projecting .................. 19

19. AR 0.26, tergite IX partly semitransparent, gonocoxite without a long dark apodeme ................................................. .............................................................................................................................. T. yakysetea Sasa et Suzuki, 2000 a

- AR 0.40–0.66, tergite IX not partly semitransparent, gonocoxite with a long apodeme ............................................... .............................................................................................................................. T. tusimufegea Sasa et Suzuki, 1999

20. Superior volsella distinct from inferior volsella ......................................................................................................... 21

- Superior volsella partly overlapping or joined with inferior volsella......................................................................... 33

21. Inferior volsella bare, with apodeme; transverse sternapodeme narrow and very thin, without oral projection............ ...................................................................................................................................... T. trivittata Goetghebuer, 1935

- Inferior volsella with thin glandular setae, without apodeme; transverse sternapodeme wide and thick, with oral pro- jection.......................................................................................................................................................................... 22

22. Inferior volsella low and placed caudally on gonocoxite ........................................................................................... 23

- Inferior volsella high and placed basally or medially on gonocoxite ......................................................................... 30

23. Third palpomere with small apical projection; gonostylus with 2 megasetae T. taurocapita Hestenes & Saether, 2000

– Third palpomere without small apical projection; gonostylus with 1 megaseta......................................................... 24

24. Superior volsella large, triangular and close to inferior volsella, conspicuously projecting ..................................... 25

- Superior volsella small, triangular or narrow and at a distance from inferior volsella, not projecting ...................... 27

25. AR about 0.30; inferior volsella with rounded corner, posterior margin of superior volsella flat ................................ ..................................................................................................................................................... T. safi Lehmann 1981

- AR about 0.50; inferior volsella caudally hooked, posterior margin of superior volsella oblique............................. 26

26. Tergite IX partly semitransparent; phallapodeme slightly curved, gonostylus narrowed near apex ............................. ............................................................................................................................ T. ginzanquerea Sasa et Suzuki, 2001

- Tergite IX not partly semitransparent; phallapodeme distinctly curved distinctly; gonostylus equally wide to apex .. ..................................................................................................................... T. oyabedilata Sasa, Kawai et Ueno, 1988

27. All trochanters of legs with dorsal keel ........................................................................................... T. liae Paggi 2007

- Trochanters without keel or only fore trochanter with dorsal keel............................................................................. 28

28. Superior volsella joined medially; inferior volsella largely typical triangular ................................... T. sichuana sp. n.

- Superior volsella separate medially; inferior volsella not largely triangular.............................................................. 29

29. Posterior of inferior volsella hooked; tergite IX not partly semitransparent .................................................................. ........................................................................................................... T. ginzanquinta ( Sasa et Suzuki, 1998) comb. n.

- Posterior of inferior volsella rounded; tergite IX partly semitransparent............................... T. spreta ( Roback, 1962)

30. Inferior volsella rectangular........................................................................................................................................ 31

– Inferior volsella largely triangular .............................................................................................................................. 32

31. Antenna with 10 flagellomeres; superior volsella conspicuously projecting ........ T. boltoni Hestenes & Saether, 2000

- Antenna with 12–13 flagellomeres; superior volsella not projecting .................................. T. minuscule Brundin 1949

32. Superior volsella triangular and projecting; phallapodeme without projection, placed below sternapodeme .............. .......................................................................................................................................................... T. wuyiensis sp. n.

- Superior volsella narrow and not projecting; phallapodeme with projection for joint with sternapodeme placed pre- lateral ................................................................................................................................................. T. triangula sp. n.

33. Antenna with 10 flagellomeres, apical part of superior volsella hooked.................................................................... 34

– Antenna with 11–13 flagellomeres, apical part of superior volsella rounded or flat.................................................. 35

34. AR 0.30; superior volsella long triangular and projecting; tergite IX not partly semitransparent ................................ ...................................................................................................................................... T. chuzeduodecima Sasa, 1984

- AR 0.46; superior volsella narrow and not projecting; tergite IX partly semitransparent.............................................. ............................................................................................................................... T. tusimuefea Sasa et Suzuki, 1999

35. Inferior volsella triangular .......................................................................................................................................... 36

- Inferior volsella nearly rectangular............................................................................................................................. 37

36. Antenna with 11 flagellomeres; AR 0.35, superior volsella projecting, triangular ........................................................ .......................................................................................................................... T. caspersi Saether et Ferrington, 2003

- Antenna with 13 flagellomeres; AR 0.27, superior volsella fused (not distinctly visible)............................................. .................................................................................................................... T. ogasaquindecima Sasa et Suzuki, 1997

37. Third palpomere with small apical projection; phallapodeme expanded medially, superior volsella nearly rectangular .......................................................................................................... T. tiunovae Makarchenko et Makarchenko 2006

- Third palpomere without apical projection; phallapodeme not expanded medially, superior volsella nearly triangular or rounded ................................................................................................................................................................... 38

38. Superior volsella separated medially, triangular, apical part nearly right-angled T. tonewquerea Sasa et Tanaka, 2002

- Superior volsella joined medially, rounded or narrowed, apical part with rounded corner........................................ 39

39. Antenna with 10 or 11 flagellomeres ......................................................................................................................... 40

- Antenna with 12 or 13 flagellomeres.......................................................................................................................... 41

40. AR 0.30–0.37; superior volsella small and not projecting; phallapodeme long and strongly curved; apex of gonosty- lus straight ......................................................................................................................... T. clavicornis Kieffer, 1911

- AR 0.60–0.76; superior volsella large and projecting; phallapodeme short and slightly curved; apex of gonostylus slightly hooked ..................................................................................................................... T. obscura Brundin, 1947

41. Superior volsella small and not projecting ................................................................................................................. 42

- Superior volsella large rounded and conspicuously projecting .................................................................................. 45

42. Anal vein of wing surpassing cubital fork; inferior volsella almost flat posteriorly .................................................. 43

- Anal vein of wing not reaching cubital fork; inferior volsella slightly oblique posteriorly ...................................... 44

43. Antenna with 13 flagellomeres; AR more than 0.50 ......................................................... T. acuticornis Kieffer, 1912

- Antenna with 12 flagellomeres; AR 0.30 ................................................... T. ogasaquardecima Sasa et Suzuki, 1997

44. Ultimate flagellomere of antenna longer than combined length of preceding eight flagellomeres; AR about 0.85 ...... ........................................................................................................................................ T. majuscula ( Edwards, 1924)

- Ultimate flagellomere of antenna shorter than combined length of preceding eight flagellomeres; AR about 0.65 ..... ............................................................................................................................................... T. lutea ( Edwards, 1924)

45. Body color very pale, AR more than 0.60 ............................................................ T. gotopallida Sasa et Suzuki, 2000

- Body color relatively dark, AR usually about 0.50 .................................................................................................... 46

46. Fore trochanter with keel, inferior volsella angle-like near apex .............................................. T. xena ( Roback, 1957)

- Fore trochanter without keel, inferior volsella rounded near apex ....................................... T. vittata ( Edwards, 1924)