Agra phallica Erwin

Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), Zootaxa 119, pp. 1-68 : 22-24

publication ID 10.5281/zenodo.162803

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Agra phallica Erwin

new species

Agra phallica Erwin , new species

( Figs. 5 a, 5 View FIGURE 5. A b, 5 c, 5 d)

Holotype. Male. COSTA RICA: Cartago, Tucurrique, 09º 51 'N 83 º 43 'W, (P. Schild & Burgdorf)( USNM: ADP 10016).

Diagnosis. Agra phallica is the only species in Costa Rica whose adults have the following attributes: (See image) Femur concolorous; elytron with sutural apex prolonged, acutely pointed (see figure). Male middle and posterior tibiae excavate, markedly arcuate, and densely setose in apical half; elytral apex shallowly arcuate between lateral and sutural apices; metasternum not pubescent; abdominal sterna 2­5 with 2 bilateral patches.

Description. Color and luster: Piceous with bright rufous appendages, head rufinistic, femur concolorous; surface very shiny. Form: Head ( Fig. 5 a View FIGURE 5. A ) behind eye moderately short, tapered in male, rounded in female. Male middle and posterior tibiae excavate, markedly arcuate, and densely setose in apical half. Elytron ( Fig. 5 View FIGURE 5. A b) with apex shallowly arcuate between lateral and sutural apices, lateral tooth small, sutural tooth acute, broad and moderately prolonged. Metasternum not pubescent. Abdominal sterna IV and V with 2 bilateral patches of dense long setae. Abdominal sternum VI ( Fig. 5 View FIGURE 5. A c) deeply round­notched in male, V­notched in female. Aedeagus ( Fig. 5 View FIGURE 5. A d) with markedly short ostium, apex a simple lobe, phallus ventrally with a longitudinal depression. Size: medium, 15.0 to 17.0mm in length, 3.40 to 4.20mm in width.

Other specimens examined. Costa Rica: 7 paratypes, 2 male, 7 females, from the following Conservation Area: Cordillera Volcánica Central. Costa Rica, Cartago, 1 f, Monumento Nacional Guayabo, Turrialba , 1100m, LN 217200,570300, June, (G. Fonseca)( INBIO : CRI 001­ 883189), 1 f, Turrialba , 3.0 km SE, CATIE area, 1100m, 83 ° 38 'W, 9 ° 53 'N, February, (H.F. Howden & A. Howden)( USNM: ADP 54792), 1 f, June, (J.E. Wappes)( USNM: ADP 58562), 1 f, April, (E. Giesbert)( CAS: ADP 62296), 1m, April, (E. Giesbert)( FSCA: ADP 103111), 1 f, (M. de Mathan)( MNHP: ADP 4859), 1 f, Tucurrique, 83 ° 43 'W, 9 ° 51 'N, (P. Schild & Burgdorf)( USNM: ADP 10015), 1m, (WAR: ADP 103113).

Specific epithet. The specific epithet, phallica , referring to the unique aedeagus, is a manuscript name from Liebke's collection in WAR. I am retaining this name, but I have chosen a different Holotype specimen than he labeled with his manuscript name, one that is in better condition.

Notes. This species is presently known from only higher altitudes, in Costa Rican wet forest.


Smithsonian Institution, National Museum of Natural History


Universidade do Extremo Sul Catarinense, Bairro Universitário


Tropical Agricultural Research and Training Center (CATIE)


California Academy of Sciences


Florida State Collection of Arthropods, The Museum of Entomology


Princeton University