Agra winnie Erwin

Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), Zootaxa 119, pp. 1-68 : 17-18

publication ID 10.5281/zenodo.162803

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Agra winnie Erwin

new species

Agra winnie Erwin , new species

( Figs. 2 a, 2 View FIGURE 2. A b, 2 c, 2 d)

Holotype. Male, COSTA RICA: Guanacaste, Guanacaste National Park, Estacíon Santa Rosa, 300m, LN 313000, 359800, (D.H. Janzen & W. Hallwachs)( INBio : CRI 000­ 590299).

Diagnosis. Apex of elytron clearly prolonged ( Fig. 2 View FIGURE 2. A b) and legs piceous with black knees.

Description. Color and luster: Piceo­castaneous, head and knees piceous; head, prontoum, and elytra very shiny. Form: Head ( Fig. 2 a View FIGURE 2. A ) behind eyes somewhat tapered in both sexes. Elytron ( Fig. 2 View FIGURE 2. A b) with apex moderately produced medially, lateral tooth small, sutural angle acute, not projected acutely; interneurs of biserite rows of punctulae. Middle tibia of male with dense brush of setae on medial margin AND metasternum pubescent, vestiture short; sternum VI ( Fig. 2 View FIGURE 2. A c) of male and female emarginate. Aedeagus ( Fig. 2 View FIGURE 2. A d) with ostium elongate; apex planariform. Stylomere 2 as in Agra duplicata (see INBio web site). Size: medium, 13.9 to 14.8mm in length, 3.8 to 4.0mm in width.

Other specimens examined. Costa Rica: 1 female paratype Specimens examined: Guanacaste, 1 f, P.N. Santa Rosa, Estacíon Santa Rosa, 300m, 85 ° 37 'W 10 ° 50 'N, LN 313000,359800, June, (D.H. Janzen & W. Hallwachs)( INBIO : CRI 000­ 590299), 1 f, Estacíon Murcielago, Cuajiniquil, 8.0 km SE, 100m, 85 ° 42 'W 10 °04'N, LN 320300,347200, July, (III curso Parataxonomos)( INBIO : CRI 000­ 927870). In addition, 2 paratypes, 1 male, 1 female, from San Salvador, El Salvador.

Specific epithet. The specific epithet, winnie , is a noun in apposition based on the given name of Winnie Hallwachs, co­collector of the Holotype.

Notes. Members of this species are similar to the preceding, however, Agra winnie is the only species in Costa Rica whose adults have all of the following attributes: elytron with convex intervals, apex clearly prolonged, legs piceous with black knees, and head somewhat tapered behind eyes.


National Biodiversity Institute, Costa Rica


Universidade do Extremo Sul Catarinense, Bairro Universitário