Agra zumbado Erwin

Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), Zootaxa 119, pp. 1-68 : 63-64

publication ID 10.5281/zenodo.162803

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Agra zumbado Erwin

new species

Agra zumbado Erwin , new species

( Figs. 30 a, 30 View FIGURE 30. A b, 30 c, 30 d)

Holotype. Male, COSTA RICA: Guanacaste, Guanacaste National Park, Estacíon Pitilla , 9.0 km S Santa Cecilia, 700m, 10 ° 59 ' 26 "N 85 ° 25 ' 40 "W, April, (C. Moraga & P. Rios)( INBio : CRI 000­ 166003).

Diagnosis. This is the only species of Costa Rican Agra with ferrugineus venter and appendages and bright brassy elytra and tri­dentate elytral apex.

Description. Color and luster: With ferruginous venter and appendages and bright brassy elytra. Form: Head ( Fig. 30 a View FIGURE 30. A ) behind eyes short, quadrate with broadly rounded hind angles. Elytron with striatio­punctate interneurs, 2, 4, and 6 with setigerous foveae; apex ( Fig. 30 View FIGURE 30. A b) slightly oblique, lateral tooth small, sutural apex rounded, not projected acutely, margin slightly lobed at middle. Male metasternum, abdominal sterna 2­6, and trochanters densely pubescent. Middle tibia of male with sparse medial brush. Sternum VI ( Fig. 30 View FIGURE 30. A c) of male shallowly V­shaped with acute posterio­lateral angles, that of female deeply notched medially with obtuse posterio­lateral angles. Aedeagus ( Fig. 30 View FIGURE 30. A d) with ostium elongate; apex of aedeagus a broad medieval axe­shaped lobe, parameres glabrous. Size: Small: 11.0 to 12.0mm in length, 3.48 to 3.62mm in width.

Other specimens examined. Costa Rica: 3 paratypes, 1 female, 2 males, from the Guanacaste Conservation Area. Costa Rica, Guanacaste, 1m, March, (C. Moraga & R. Blanco)( INBIO : CRI 000­ 211532), 1 f, March ­ April, (C. Moraga)( INBIO : CRI 000­ 748607), 1m, May, (P. Rios)( INBIO : CRI 001­ 877475).

Specific epithet. The specific epithet, zumbado , is used as a noun in apposition based on the family name of the collector of many interesting Agra specimens, ex­parataxonomist, and now INBio Inventory chief, Manuel A. Zumbado of San Antonio de Belén, Heredia, Costa Rica.

Notes. This species features all the external attributes of typical purpurea group members, however, I have classified it in its own subgroup because of the shorter stylomere 2 and the broad­ax shape of the aedeagal apex.


National Biodiversity Institute, Costa Rica


Universidade do Extremo Sul Catarinense, Bairro Universitário