Stenaesthetus SHARP

Puthz, V., 2013, Revision der orientalischen Stenaesthetus-Arten (Coleoptera: Euaesthetinae) 114. Beitrag zur Kenntnis der Euaesthetinen, Linzer biologische Beiträge 45 (2), pp. 2077-2113 : 2101-2103

Key to the Oriental species of the genus Stenaesthetus SHARP

(FL = furrow length, S = sternite, ST = spermatheca)

1 Pronotum simple, without longitudinal furrows or lateral rugae (figs 1, 2)........................2

- Pronotum otherwise; with longitudinal furrows and/or lateral rugae (figs 3-9)..................5

2 Elytra longer (EW: EL =1.09-1.30), shoulders simple, abdominal sculpture scabrous.......3

- Elytra shorter ( EW: EL =131-140, Ø 1.35; fig. 1). Shoulders with a short carina, abdomen punctate.: S 8 (as in fig. 26). E (fig. 57). 2.2-2.8mm.: ST (fig. 101). (FB 1.2-1.3mm). India (Meghalaya) ........................................................... St. scabratus nov.sp .

3 Elytra longer (EW: EL =1.09-1.26)....................................................................................4

- Elytra less long (EW: EL =1.22-1.21, Ø 1.27): S 8 (fig. 26). E (fig. 56). 2.2-2.8mm (FB 1.2-1.3mm). India (Assam, W. Bengal, Arunachal Pradesh) ............. St. scaber nov.sp.

4 Larger: 2.7-3.3mm (FB 1.5-1.6mm), head narrower (HW: EW Ø 0.80).: Excision of S 8 longer than half the length of S (fig. 35). S 9 (fig. 30). E (fig. 58).: ST (fig. 100). China (Yunnan), Thailand.................................................................. St. conflictatus PUTHZ

- Smaller: 1.9-2.5mm (FB 1.1-1.2mm), head broader (HW: EW Ø 0.85).: Excision of S 8 less long than half the length of S (fig. 34). S 9 with some dorsal setae. E (fig. 55).: ST (fig. 102). Japan, China (Sichuan, Yunnan, Taiwan, Shanghai, Guangxi, Beijing), Vietnam, Thailand, Nepal, India, Pakistan, Sri Lanka, Indonesia (Sulawesi) ........ ............................................................................................................... St.sunioides SHARP

5 Pronotum with a short, very narrow lateral carina..............................................................6

- Pronotum with a long lateral furrow...................................................................................8

6 Punctation of pronotum slightly rugose, mostly distinctly delimited. Larger species.........7

- Sculpture of pronotum rugose longitudinally, punctures in long and dense rugae. Smaller: 1.8-2.1mm (FB 0.8-1.0mm).: S 8 (fig. 47). E (fig. 59). Vietnam....................... ................................................................................................................. St. perroti nov.sp.

7: S 7 (fig. 23), S 8 (fig. 37). E (fig. 33).: ST (fig. 93). 1.9-2.6mm (FB 1.0-1.2mm). Thailand, Vietnam, China (Sichuan, Yunnan, Guizhou, Guangxi, Hongkong) ..................... ..................................................................................................... St. deharvengi OROUSSET

-: S 7 (fig. 24), S 8 (fig. 41). E (fig. 29).: ST (as in fig. 93). 1.8-2.2mm (FB 0.9- 1.0mm). Thailand.................................................................................... St. odiosus nov.sp.

8 Pronotum, shiny, impunctate or shallowly punctate............................................................9

- Pronotum matt or slightly shiny, densely punctate/sculptured..........................................10

9 Smaller, frons reticulate, labrum pointed.: S 8 (fig. 46), E (fig. 60).: Unknown. 1.8mm (FB (0.8mm). Myanmar................................................................... St. naso nov.sp.

- Larger, frons punctate, interstices smooth, labrum simple.: S 8 (fig. 39). E (fig. 61).: ST (fig. 92). 2.0-2.4mm (FB 1.1-1.2mm). India (Assam)............. St. nitidicollis nov.sp.

10 Frons and clypeus uniformly sculptured...........................................................................16

- Frons and clypeus differently sculptured, clypeus ± set off from frons............................11

11 Smaller, FB <0.8mm.: S 8 (fig. 42). E (fig. 63). 1.7-1.9mm (FB 0.8mm). Malaysia........ ........................................................................................................ St. frontalis (CAMERON)

- Larger, FB 1.0mm...........................................................................................................12

12 Distance between longitudinal furrows of pronotal disc from anterior margin of pronotum less than length of antennal segment 3..............................................................13

- Distance between longitudinal furrows of pronotal disc from anterior margin of pronotum wider, at least as wide as length of antennal segment 3....................................15

13: E (figs 26-28). 2.3-2.8mm (FB 1.0-1.2mm). Nepal, India (Sikkim, W. Bengal) ............. ............................................................................. St. canaliculatus canaliculatus COIFFAIT

-: E (fig. 32). 2.4-2.9mm (FB 1.2mm). Nepal................. St. canaliculatus taurulus PUTHZ

14 Frons moderately shiny, densely sculptured with shallow longitudinal rugae (fig. 3). Larger: 2.6-3.0mm (FB 1.3-1.6mm).: E (fig. 65).: ST (fig. 96). China (Yunnan) ........ ............................................................................................................ St. hastipenis nov.sp.

- Frons shiny, nearly smooth. Smaller.................................................................................15

15: S 8 (fig. 51). E (fig. 66).: ST (fig. 91). 2.1-2.7mm (FB 1.1-1.2mm). Nepal................ ............................................................................................................. St. nitidifrons PUTHZ

-: E (fig. 62). 1.9-2.4mm (FB 1.0-1.1mm). India (Darjeeling distr.) ................................... ....................................................................................................... St. angulatipenis nov.sp.

16 Longitudinal furrows of pronotal disc shorter, less long than half the length of .............the pronotum (e. g. figs 8, 9)...................................................................................................36

- Longitudinal furrows of pronotal disc longer than half the length of the pronotum (e. g. figs 3-7)....................................................................................................................17

17 Longitudinal furrows of pronotal disc broader, as broad as antennal segment 2...............18

- Longitudinal furrows of pronotal disc less broad, narrower than antennal segment 2......20

18 Pronotum deply and densely striate longitudinally, labrum simple.: S (fig. 49). E (fig. 67). 1,8mm (FB 0.8mm). Thailand........................................... St. rimosicollis nov.sp.

- Pronotum deeply and densely punctate (fig. 7), labrum with two tiny median calli. 19

19: S 8 (fig. 45). E (fig. 88). 1.5-1.9mm (FB 0.8mm). Malaysia......... St. malayanus PUTHZ

-: S 8 (as in fig. 45). E (fig. 68). 1.5-1.9mm (FB 0.8mm). Malaysia....... St. persimilis nov.sp.

20 Longitudinal furrows of pronotal disc longer, distance from anterior margin of pronotum distinctly less than one eye-length....................................................................21

- Longitudinal furrows of pronotal disc about as long as one eye-length or longer.............25

21 Elytra less broad (EW: EL <1.40). Paraglossae prominent, spine-shaped (fig. 13)..........22

- Elytra broader (EW: EL>1.40). Paraglossae not prominent, not spine-shaped (figs 10- 12) .....................................................................................................................................23

22 Elytra deeply and densely punctate.: S 8 (fig. 52). E (fig. 69). 2.3mm (FB 1.2mm). China (Yunnan).................................................................................... St. schuelkei nov.sp.

- Elytra less deeply, moderately densely punctate.: S 8 (fig. 43). E (fig. 73). 2.5mm (FB 1.3mm). China (Yunnan) ............................................... St. gaoligongmontium nov.sp.

23 Furrows of pronotal disc nearly attaining anterior margin of pronotum. Sculpture of pronotum densely reticulate, no distinct punctation.: E (fig. 70).: ST (fig. 102). 1.0-2.0mm (FB 1.0mm). Nepal............................................................. St. smetanai nov.sp.

- Furrows of pronotal disc distinctly separate from anterior margin of pronotum. Anterior pronotum distinctly punctate..............................................................................24

24: S 8 (fig. 44). E (figs 74, 75). Punctation of pronotum less deep. 1.9-2.2mm (FB 1.0- 1.1mm). India (West Bengal) .................................................................. St. hastifer nov.sp.

-: E: (fig. 76). 1.9- 2,3mm (FB 1.0-1.1mm). India (W. Bengal)....... St. indicus (CAMERON)

25 Frons impunctate...............................................................................................................26

- Frons ± distinctly punctate................................................................................................29

26 Pronotum impunctate........................................................................................................27

- Pronotum on anterior half ± distinctly punctate, punctures elongated longitudinally.: E (fig. 71). 2.0-2.5mm (FB 1.0mm) .................................................... St. nepalensis PUTHZ

27: E (fig. 72). 2.0-2.4mm (FB 1.0-1.1mm). Nepal................................ St. martensi PUTHZ

-: otherwise......................................................................................................................28

28: E (fig. 64). 2.3-2.8mm (FB 1.0-1.2mm). Nepal, India (Sikkim, W. Bengal) ................... ................................................................................................ St. quadrisulcatus CAMERON

-: S 8 (fig. 48). E (fig. 77). 2.0-2.3mm (FB 1.1mm). China (Yunnan) .......... St. puetzi nov.sp.

29 Pronotum coarsely and densely punctate, punctures distinctly delimited, as large as apical cross section of antennal segment 2, interstices smooth, shiny.: E (fig. 83). 2.0-2.4mm (FB 1.1mm). Laos.............................................................. St. kurbatovi nov.sp.

- Sculpture of pronotum otherwise (punctures less large/elongate longitudinally/interstices reticulate)..................................................................................30

30 Species from Taiwan.........................................................................................................31

- Species from the Asian mainland......................................................................................33

31: E (fig. 78).: ST (fig. 11, PUTHZ 1995) y). 2.0-2.2mm (FB 1.0mm). China (Taiwan) .................................................................................................. St. nomurai PUTHZ

-, otherwise.................................................................................................................32

32: Sides of posterior impression of S 6 slightly prolonged posteriad, posterior margin broadly emarginate, posterior margin of S 7 slightly emarginate medially. E (fig. 80).: ST (fig. 95). 2.0-2.6mm (FB 1.1mm) China (Taiwan) ................. St. formosanus nov.sp.

-: Sides of S 6 not prolonged posteriad, posterior margin broadly emarginate, posterior margin of S 7 simple. E (fig. 79). 2.0-2.3mm (FB 1.0-1.1mm). China (Taiwan) ............................................................................................. St. taiwanensis PUTHZ

33 Posterior middle of pronotum between medial disc furrows deeply and densely sculptured.: S 8 (as in fig. 54).: ST (about as in fig. 94). 1.9-2.4mm (FB 1.0- 1.1mm). Thailand.......................................................................................... St. thai nov.sp.

- Posterior middle of pronotum between medial disc furrows less deeply and less densely sculptured than anterior mddle of pronotum........................................................34

34 Lateral portion of pronotum beside lateral longitudinal furrow narrowly not smooth but finely sculptured/punctate.................................................................................................35

- Lateral portion of pronotum beside lateral longitudinal furrow completely impunctate/smooth: see # 27 ..................................................... St. quadrisulcatus COIFFAIT

35: S 8 (fig. 40). E (fig. 86).: ST (fig 20). 1.8-2.5mm (FB 1.1mm). Thailand................... ................................................................................................................... St. loebli nov.sp.

-: S 8 (fig. 53). E (fig. 84).: ST (about as in fig. 94). 1.9-2.3mm (FB 1.0-1.1mm). Thailand............................................................................................... St. siamensis nov.sp.

36 Punctures of pronotum deeper, elongated longitudinally but not in long shallow rugae.: S 8 (fig. 50). E (fig. 87).: ST (fig. 98). 1.9-2.4mm (FB 1.0-1.1mm) China (Fujian, Jiangxi, Zhejiang) ............................................................. St. brevisulcatus nov.sp.

- Punctures of pronotum less deep, arranged in long shallow longitudinal furrows............37

37 Longitudinal furrows of the pronotal disc shorter (PL: FL =52: 21/ 47: 21).: E (fig. 82).: ST (fig. 89). Japan (Iriomote Is., Ishigaki Is.)........................... St. insulanus PUTHZ

- Longitudinal furrows of the pronotal disc longer..............................................................38

38 Head narrower than elytra (HW: EW <0,95). PL: FL =49: 29/ 51: 31).: E (fig. 81).: ST (fig. 90). 2.0-2.4mm (FB 1.0-1.1mm) Japan (Ie Rindo, Kuweijima Is., Amami-Oshima Is., Amami Is.) ................................................................... St. okinawaensis PUTHZ

- Head as broad as elytra (HW: EW = 1.0). PL: FL =51: 28).: Unknown.: ST (fig. 99). 2.0mm (FB 1.05mm). Japan (Amami Oshima Isl.).......................... St. lackneri PUTHZ