Cochlostoma (T.) mariannae Nordsieck, 2011

Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De & Gittenberger, Edmund, 2024, Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae), European Journal of Taxonomy 927, pp. 1-163 : 107-108

Cochlostoma (T.) mariannae Nordsieck, 2011

Figs 82 (yellow dots), 95–96

Cochlostoma mariannae Nordsieck, 2011: 15–16 .

Types

Not seen.

Other specimens

ITALY • 1- Vado di Sole 1(topotypical); 42.3968° N, 13.788° E; 2011; Hallgass leg.; EZ1084 • 2- Vado Di Sole 2 (topotypical); 42.3968° N, 13.788° E; 2013; Margelli leg.; EZ1145 • 3- Rigopiano ; 42.43° N, 13.7807° E; 2013; Margelli leg.; EZ1140 • 4- Farindola loc. Angri ; 42.4442° N, 13.8199° E; 2013; Margelli leg.; EZ1141 • 5- Rigopiano prov 37 ; 42.4482° N, 13.7501° E; 2013; Margelli leg.; EZ1143 • 6- Rigopiano camping ; 42.4306° N, 13.7795° E; 2013; Margelli leg.; EZ1144 .

Type locality

ITALY • L’Aquila , Vado di Sole .

Description

SHELL. Closely spaced riblets on last part of protonch. Teleoconch with two barely visible lines of reddish spots on whorls. Moderately strong, often whitish ribs becoming weaker approaching aperture. Welldeveloped and strong lip with slightly and gently curved columellar lobe almost completely covering umbilicus.

MEASUREMENTS. 11 ♀♀: whorls=7.2–8.4, H= 7.3–9.8 mm, H/W= 2.39–2.83, roundness =0.11–0.17, ribs incl.= 55–62°, apert. incl. =16–25°, ribs/mm 1 st wh. =6–16, ribs/mm 4 th wh.=5–9.

FEMALE GENITAL ORGANS. As in C. (T.) nanum .

Remarks

Although this species was described recently (2011), only the shell morphology was taken into consideration in the original description.

Additional note to clade B

The following species C. (T.) affine (Benoit, 1879) , C. (T.) agriotes ( Westerlund, 1879) and C. (T.) crosseanum ( Paulucci, 1879) have an almost identical shell and female genital morphology. Despite the morphological similarity, only agriotes and crosseanum cluster together whereas affine appears in an unresolved politomy. They live in the central and northern parts of the Apennines and in Sicily.

Figures

Fig. 82.Distributionof samplesin Clade B: red= Cochlostoma (T.)braueri (Wagner, 1897);purple=C. (T.) stossichi (Hirc, 1881); blue=C. (T.) hallgassi sp. nov.; yellow =C. (T.) mariannae Nordsieck, 2011; green =C. (T.) dubium (Wagner, 1897); white = C. (T.) mostarense (Wagner, 1906). Black center for the samples with amplified 16S.