Scrapter nitidus (Friese, 1909)

Mack, Anne & Kuhlmann, Michael, 2023, Revision of the nitidus species group of the bee genus Scrapter Lepeletier & Serville, 1828 (Hymenoptera: Apoidea), European Journal of Taxonomy 912, pp. 1-119 : 112-117

publication ID	https://doi.org/ 10.5852/ejt.2023.912.2373
publication LSID	lsid:zoobank.org:pub:B1958762-5D89-4DC9-A8CE-AABB06B7979C
DOI	https://doi.org/10.5281/zenodo.10420150
persistent identifier	https://treatment.plazi.org/id/03A4762C-FFD5-CA2C-FF71-352AFAB4D36A
treatment provided by	Plazi
scientific name	Scrapter nitidus
status

Treatment

Key for the identification of species in the Scrapter nitidus View in CoL species group

Females

Females of S. convexoides sp. nov., S. perpunctatulus sp. nov. and S. glaberrimus are unknown.

1. Pronotal lobe yellowish to white ( Figs 2A View Fig , 3A View Fig ); if pronotal lobe brownish, scutellum and metanotum at least partly orange ( Figs 70C View Fig , 73C View Fig ) ............................................................................................... 2

– Pronotal lobe brownish to black ( Figs 24A View Fig , 36A View Fig , 57A View Fig ); scutellum and metanotum dark brown to black................................................................................................................................................... 6

2. Scutum, scutellum and metanotum partly orange; propodeum completely rugulose-areolate; supraclypeal area delimited by sharp edges ( Figs 70C View Fig , 73C View Fig )............................................................ 3

– Scutum, scutellum and metanotum brownish or black; propodeum often partly carinate or coriaceous ( Figs 8C–D View Fig , 58C–D View Fig , 64C–D View Fig ); supraclypeal area elevated but not sharply delimited....................... 4

3. Metanotum with an orange spot medially ( Fig. 70C View Fig ).................................. S. flavipunctatus sp. nov.

– Metanotum completely brown ( Fig. 73C View Fig ) .......................................... S. ruficornis ( Cockerell, 1916)

4. Facial fovea narrow, at least 5 times as long as wide, convergent at vertex ( Fig. 8B View Fig ); posterior ocelli in line with posterior margin of compound eyes; discs of T2–T3 deeply punctate ( Fig. 8E–F View Fig ) ......... ............................................................................................................ S. flavostictus Cockerell, 1934 View in CoL

– Facial fovea wide, about 3.5 times as long as wide ( Fig. 3B View Fig ), parallel at vertex; posterior ocelli in front of posterior margin of compound eyes; discs of T1–T3 shallowly punctate ( Figs 2E–F View Fig , 21E– F View Fig )........................................................................................................................................................ 5

5. Scutum sparsely punctate (i = 1–2.5 d) ( Fig. 2C View Fig )............................. S. basutorum ( Cockerell, 1915) View in CoL

– Scutum densely punctate (i = 0.1 d) ( Fig. 5C View Fig )............................................. S. flavipes ( Friese, 1925) View in CoL

6. Surface of scutum between coarse punctation with fine punctation ( Figs 39C View Fig , 78C View Fig )....................... 7

– Scutum only with coarse punctation ( Figs 27C View Fig , 54D View Fig , 61C View Fig ).............................................................. 8

7. Discs of T1–T3 with very fine and coarse punctation (i = 1–2 d) ( Fig. 76E View Fig ); facial fovea about 4 times as long as wide ( Fig. 76B View Fig ) ......................................................................... S. confusus sp. nov.

– Discs of T1–T3 almost impunctate ( Fig. 39E View Fig ); facial fovea about 8 times as long as wide ( Fig. 39B View Fig )............................................................................................................... S. obtusus sp. nov.

8. T3 anterior of premarginal line without row of punctures ( Figs 21E View Fig , 33E View Fig ); if T3 is completely punctate, punctation is regular and deep, single points are separated ( Fig. 58E View Fig ).............................. 9

– T3 anterior of premarginal line with a row of punctures ( Figs 45D View Fig , 55C–D View Fig ); if T3 is completely punctate, punctation is irregular, single points merge into each other............................................. 18

9. Supraclypeal area at upper margin straight to slightly convex ( Figs 27B View Fig , 58B View Fig , 61B View Fig ); propodeum shallowly carinate, at least partly smooth or coriaceous ( Figs 27D View Fig , 58D View Fig , 61D View Fig ) (It is assumed that S. flavitarsis belongs to this group based on the characteristics of similar species and S. cf. flavitarsis ) ........................................................................................................................................ 10

– Supraclypeal area at upper margin distinctly emarginated and edges slightly extended ( Figs 24B View Fig , 48B View Fig , 51B View Fig ); propodeum usually more deeply carinate or areolate, sometimes with smooth parts laterally ( Figs 24D View Fig , 48D View Fig , 51D View Fig ) ........................................................................................................ 16

10. Propodeum coriaceous on posterior half ( Fig. 21D View Fig ); scutum densely (i = 0.5 d) punctate ( Fig. 21C View Fig ); T2–T3 superficially and regularly (i ~ 2 d) punctate; scopa greyish to brownish, without black hair ( Fig. 21A View Fig ); supraclypeal area distinctly elevated, formed like pyramid ( Fig. 21B View Fig )............................ ............................................................................................................... S. flavitarsis Cockerell, 1936

– Propodeum smooth and shiny on posterior half ( Figs 27D View Fig , 48D View Fig , 58D View Fig ); scutum sparsely punctate (i> 0.5 d) ( Figs 27C View Fig , 48C View Fig , 58C View Fig ); T2–T3 either less punctate or more deeply; scopa often partly black ( Fig. 48A View Fig ); supraclypeal area usually flat to slightly elevated ( Figs 27B View Fig , 48B View Fig , 58B View Fig )......................11

11. Discs of T1–T4 deeply and regularly punctate ( Figs 58E View Fig , 61E View Fig , 64E View Fig ); posterior ocelli in line with posterior margin of compound eyes................................................................................................. 12

– Discs of T1–T4 shallowly punctate ( Figs 33E View Fig , 48E View Fig ); posterior ocelli in front of posterior margin of compound eyes ( Fig. 11D View Fig )............................................................................................................... 14

12. Disc of T2 basally denser (i = 0.5–1 d) punctate than apically (i = 2–3 d) ( Fig. 58E View Fig ). Scutum densely punctate, laterally denser (i = 0.5 d) than medially (i = 1–1.5 d) ( Fig. 58C View Fig ); propodeum smooth and shiny ( Fig. 58D View Fig ); supraclypeal area elevated, but not sharply delimited ( Fig. 58B View Fig )........................... ................................................................................................................. S. divergens ( Friese, 1925)

– Disc of T2 basally denser (i = 1–2 d) punctate than apically (i = 3–5 d) ( Figs 61F View Fig , 64F View Fig ); scutum completely irregularly (i = 0.5–2 d or 2–4 d) punctate ( Figs 61C View Fig , 64F View Fig ); propodeum matt ( Figs 61D View Fig , 64F View Fig ); supraclypeal area elevated and sharply delimited ( Figs 61B View Fig , 64F View Fig ) ....................................... 13

13. Scutum finely and densely (i = 0.5–2 d) punctate ( Fig. 61C View Fig )............................ S. montanus sp. nov.

– Scutum coarsely and sparsely (i = 2–4 d) punctate ( Fig. 64C View Fig )....... S. pallidicinctus Cockerell, 1933

14. Prepygidial and pygidial fimbria and scopa partly black ( Fig. 27A View Fig ); propodeum basally with fine, v-shaped carinae, apically smooth and shiny ( Fig. 27D View Fig ); mesosoma shallowly punctate ( Fig. 27E– F View Fig ).......................................................................................................................... S. imparilis sp. nov.

– Prepygidial and pygidial fimbria at most dark brown ( Figs 12A View Fig , 33A View Fig ); propodeum different ( Figs 12B View Fig , 33D View Fig ); metasoma almost impunctate ( Figs 12C–D View Fig , 33E–F View Fig )........................................... 15

15. Propodeum basally with carinae ( Fig. 33D View Fig ); metasoma smooth but without oily-bluish shine; mesepisternum with coarse punctation mixed with fine punctation; surface between punctures matt ( Fig. 33C, E–F View Fig ) ...................................................................................... S. mpumalangensis sp. nov.

– Propodeum basally coriaceous ( Fig. 12B View Fig ); metasoma with oily-bluish shine; mesepisternum only with coarse punctation, surface between punctures smooth and shiny ( Fig. 12C–D View Fig )......................... ........................................................................................................................... S. caeruleus sp. nov.

16. T1 anterior of premarginal line with a row of punctures, disc of T1 without punctation, surface distinctly matt ( Fig. 48F View Fig ); propodeum deeply carinate ( Fig. 48D View Fig ) ............................................... .............................................................................................................. S. semirufus Cockerell, 1932

– T1 with very fine and dispersed punctures (i> 5 d) ( Figs 24F View Fig , 51F View Fig ), surface slightly matt; propodeum shallowly carinate ( Figs 24E View Fig , 51D View Fig ) ................................................................................................. 17

17. Transitional line from disc to apical depression of T1–T4 curved ( Fig. 51E–F View Fig ); discs of T1– T4 shallowly punctate, punctures almost invisible ( Fig. 51E–F View Fig ) ....... S. sphecodoides ( Friese, 1912)

– Transitional line from disc to apical depression of T1–T4 straight ( Fig. 24F View Fig ); disc of T1 shallowly punctate, discs of T2–T3 deeply punctate ( Fig. 24F View Fig ) ............................ S. fuscipennis ( Friese, 1912)

Females of the following species are very similar. Their identification can be difficult without reference specimen.

18. Discs of T1–T2 almost impunctate, sometimes punctation hardly visible, T1 anterior of premarginal line without row of punctures ( Fig. 30E–F View Fig ); disc of T3 deeply punctate; metasoma matt ( Fig. 30E View Fig ); scutum regularly and densely (i = 1–2 d) punctate ( Fig. 30 C View Fig ); transitional line between anterior longitudinal carinae and posterior transverse carinae of propodeum curved ( Fig. 30D View Fig ) .................... .............................................................................................................................. S. littoralis sp. nov.

– Discs of T1–T3 with distinct punctation, if punctation shallowly, then anterior of premarginal line of T1 with row of punctures; transitional line between anterior longitudinal and posterior transverse carinae usually straight; other combination of characters ( Figs 15C–F View Fig , 36C–F View Fig , 67C–F View Fig )................ 19

19. Discs of T1–T2 shallowly and dispersed punctate (i> 4 d), T1 anterior of premarginal line with row of punctures ( Fig. 55C–D View Fig ); propodeum finely carinate, laterally often smooth and shiny ( Fig. 55A– B View Fig ); scutum medially more densely punctate than laterally ( Fig. 54D View Fig )................ S. variabilis sp. nov.

– Discs of T1–T2 more deeply punctate; other combination of characters ( Figs 15C–F View Fig , 36C–F View Fig , 67C– F View Fig )...................................................................................................................................................... 20

20. Discs of T1–T3 finely and shallowly punctate; propodeum finely carinate ( Fig. 36D View Fig ) ...................... ...................................................................................................................... S. nitidus ( Friese, 1909) View in CoL

– Discs of T1–T3 coarsely and deeply punctate; propodeum different ( Figs 15C–F View Fig , 18C–F View Fig , 67C–F View Fig ) .. ......................................................................................................................................................... 21

21. Scutum and mesosoma regularly and densely (i = 1–2 d) punctate ( Fig. 18C View Fig ); disc of T3 basally often more densely (i <1 d) punctate than apically (i = 1–2 d) ( Fig. 18E–F View Fig ); propodeum laterally often smooth and shiny, posterior carinae shallow ( Fig. 18D View Fig ) ................. S. crassipunctatus sp. nov.

– Scutum and mesosoma more irregular and/or sparser punctate; disc of T3 usually regularly punctate; propodeum different. If basal third of disc of T3 more densely punctate than posterior third, propodeum coarsely carinate; ( Figs 15C–F View Fig , 44D View Fig , 45A–D View Fig , 67C–F View Fig ) ................................................ 22

22. Propodeum laterally carinate, medially (rugulose-)areolate ( Fig. 67D View Fig ); mesepisternum irregularly (i = 0.5–3 d) punctate; discs of T1–T4 regularly punctate (i = 3–4 d); T3 sometimes more densely (i = 2–3 d) punctate than T2 (i = 3–4 d) ( Fig. 67E–F View Fig ).................................................... S. felicis sp. nov.

– Propodeum and punctation of mesosoma different ( Figs 15C–F View Fig , 44D View Fig , 45A–D View Fig )............................. 23

23. Discs of T2–T3 regularly (i = 2–3 d) punctate ( Fig. 15E–F View Fig ); propodeum medially coarsely and irregularly areolate to longitudinally carinate, laterally carinate, posterior third transversely carinate ( Fig. 15D View Fig ); posterior ocelli little behind posterior margin of compound eyes ... S. convexus sp. nov.

– Discs of T2–T3 irregularly ( Fig. 45C–D View Fig ) punctate; propodeum coarsely carinate ( Fig. 45A–B View Fig ); posterior ocelli in line with posterior margin of compound eyes .... S. perpunctatus Cockerell, 1933

Males

Males of S. caeruleus sp. nov., S. fuscipennis and S. littoralis sp. nov. are unknown.

1. Pronotal lobe yellow to white ( Figs 3A View Fig , 7A View Fig , 10A View Fig )............................................................................ 2

– Pronotal lobe brown to black ( Figs 34A View Fig , 62A View Fig , 77A View Fig )......................................................................... 4

2. Facial fovea wide, about 3.5 times as long as wide; mesosoma covered with long hair ( Fig. 3D View Fig ); hind femur partly yellow ( Fig. 3A View Fig ).......................................................... S. basutorum ( Cockerell, 1915) View in CoL

– Facial fovea narrow, at least 5 times as long as wide; mesosoma covered with short hair ( Figs 7D View Fig , 10D View Fig ); hind femur completely dark brown to black ( Figs 7A View Fig , 10A View Fig ).................................................. 3

3. T3 anterior of premarginal line with narrow band of pilosity of long, whitish, erect hair ( Fig. 6D View Fig ); scutum very densely (i = 0.1 d) and finely punctate ( Fig. 6B View Fig ); apical plate of S8 round in shape, shaft distinctly shorter ( Fig. 7D View Fig ) ......................................................................... S. flavipes ( Friese, 1925) View in CoL

– T3 anterior of premarginal line with hair patches laterally of short, whitish, erect hair ( Fig. 9D View Fig ); scutum densely (i = 0.5 d) and coarsely punctate ( Fig. 9B View Fig ); apical plate of S8 truncate in shape, shaft distinctly longer ( Fig. 10D View Fig ) ............................................................... S. flavostictus Cockerell, 1934 View in CoL

4. Surface of scutum between coarse punctation with fine punctation ( Fig. 77B View Fig ); facial fovea about 3 times as long as wide; flagella of antennae shorter than the compound eye ( Fig. 77A View Fig ) ..................... ............................................................................................................................ S. confusus sp. nov.

– Scutum only with coarse punctation ( Figs 14B View Fig , 19B View Fig , 59B View Fig ); facial fovea at least 6 times as long as wide; flagella of antennae longer than compound eye ( Figs 14A View Fig , 19A View Fig , 59A View Fig ) .................................. 5

5. Discs of T1–T3 sparsely punctate (i>1 d), on T2–T4 sometimes more densely punctate on basal third than on posterior third ( Figs 22D View Fig , 34D View Fig , 75D View Fig )........................................................................... 6

– At least basal third of discs of T1–T3 densely (i <0.5 d) punctate ( Figs 37D View Fig , 49D View Fig , 56D View Fig )............. 10

6. Discs of T1–T3 shallowly and finely punctate ( Figs 22D View Fig , 34D View Fig ); legs and/or tarsi yellow ( Figs 22A View Fig , 34A View Fig ) ................................................................................................................................................... 7

– Discs of T1–T3 deeply punctate ( Figs 61D View Fig , 65D View Fig , 74D View Fig ); legs completely brown to black ( Figs 61A View Fig , 65A View Fig , 74A View Fig ) .......................................................................................................................................... 8

7. Scutum finely and sparsely punctate (i = 2–3 d) ( Fig. 34B View Fig ); distal margin of S7 with deeper emargination ( Fig. 35C View Fig ); posterior ocelli behind posterior margin of compound eyes....................... ............................................................................................................... S. mpumalangensis sp. nov.

– Scutum coarsely and densely (i = 1–2 d) punctate ( Fig. 22B View Fig ); distal margin of S7 with shallower emargination ( Fig. 23C View Fig ); posterior ocelli in line with the posterior margin of compound eyes.......... .............................................................................................................. S. flavitarsis Cockerell, 1936

8. Disc of T1 sparsely and regularly (i = 3–4 d) punctate ( Fig. 75A View Fig ); propodeum coarsely areolate ( Fig. 74C View Fig ); apicolateral parts of S7 slightly curved ( Fig. 75C View Fig ) .......... S. ruficornis ( Cockerell, 1916)

– Disc of T1 densely punctate (i = 1–2 d) ( Figs 62A View Fig , 66A View Fig ); propodeum different ( Figs 61C View Fig , 65C View Fig ); apicolateral parts of S7 distinctly curved ( Figs 62C View Fig , 66C View Fig )................................................................ 9

9. Disc of T1 coarsely and densely punctate (i = 1–2 d), mixed fine punctures ( Fig. 66A View Fig ); T2 basally denser (i ~ 1 d) punctate than apically (i ~ 2 d) ( Fig. 66A View Fig ); propodeum finely areolate ( Fig. 65C View Fig ) ... ....................................................................................................... S. pallidicinctus Cockerell, 1933

– Discs of T1 and T2 finely and regularly (i = 1–1.5 d) punctate ( Fig. 62A View Fig ); propodeum coarsely areolate ( Fig. 61C View Fig )............................................................................................ S. montanus sp. nov.

10. Discs of T1–T3 evenly densely (at most i = 0.5–1 d) punctate, T2–T3 anterior of premarginal line narrowly impunctate, width at most quarter of tergum ( Figs 32D View Fig , 37D View Fig ). If discs of T1–T4 without impunctate stripe ( Fig. 59D View Fig ), scutum also completely densely (i = 0.1 d) punctate ( Fig. 59B View Fig ).......11

– Discs of T1–T3 unevenly punctate, basally more densely (i <1 d) punctate than apically (i> 1 d), T1–T3 anterior of premarginal line impunctate, width at least quarter of tergum ( Figs 31D View Fig , 47D View Fig , 56D View Fig ) ....... 13

11. Scutum completely densely (i = 0.1 d) punctate ( Fig. 32B View Fig ) ( Fig. 19B View Fig ); discs of T1–T3 completely punctate ( Fig. 19D View Fig ).................................................................................. S. crassipunctatus sp. nov.

– Scutum coarsely and sparsely (i = 0.5–1 d) punctate ( Figs 37C View Fig , 60C View Fig ); T1–T3 anterior of premarginal line sometimes narrowly impunctate ( Figs 37D View Fig , 60D View Fig ).................................................................... 12

12. Discs of T1–T4 regularly and more sparsely (i = 0.5–1 d) punctate ( Fig. 59D View Fig ); distal margin of S7 with broad emargination ( Fig. 60C View Fig ) ......................................................... S. divergens ( Friese, 1925)

– Discs of T1–T4 irregularly and more densely (i <0.3 d) punctate ( Fig. 37E View Fig ); distal margin of S7 with almost no emargination ( Fig. 38B, D View Fig ) .......................................................... S. nitidus ( Friese, 1909) View in CoL

13. S7 apicolaterally with hairs ( Figs 69C View Fig , 72C View Fig )................................................................................... 14

– S7 apicolaterally without hairs ( Figs 14C View Fig , 29C View Fig , 50C View Fig )..................................................................... 15

14. T2 on basal third of disc finely and densely punctate (i = 0.5 d) ( Fig. 69A View Fig ); scutum regularly (i = 1–1.5 d) punctate (68B); fore femur and tibia mostly brownish ( Fig. 68A View Fig ) ............ S. felicis sp. nov.

– T2 on basal fifth of disc finely and more sparsely punctate (i> 1 d) ( Fig. 72A View Fig ); scutum irregularly (i = 0.5–2.5 d) punctate ( Fig. 71B View Fig ); fore femur and tibia mostly yellowish ( Fig. 68A View Fig )...................... ..................................................................................................................... S. flavipunctatus sp. nov.

15. Terga distinctly convex in lateral view ( Figs 14A View Fig , 16A View Fig )................................................................. 16

– Terga (particularly T1) slightly convex in lateral view or usually even ( Figs 28A View Fig , 52A View Fig , 56A View Fig )...... 17

16. Discs of T1–T3 with dense (i = 0.2 d) punctation that covers basal third of tergum ( Fig. 16D View Fig ); distal margin of S7 almost without emargination ( Fig. 17C View Fig ) ........................................ S. convexus sp. nov.

– Discs of T1–T3 with dense (i = 0.2 d) punctation that covers at least basal half of tergum ( Fig. 13D View Fig ); distal margin of S7 with emargination ( Fig. 14C View Fig ) .......................................... S. convexoides sp. nov.

17. Discs of T1–T3 superficially punctate ( Figs 49D View Fig , 52D View Fig ); disc of T1 finely punctate ( Figs 50A View Fig , 53A View Fig ); basomedially on T2–T3 hair bands of short and fine hair ( Figs 49D View Fig , 52D View Fig ).................................... 18

– Discs of T1–T3 deeply punctate ( Figs 25D View Fig , 27D View Fig , 31D View Fig ); disc of T1 coarsely punctate ( Figs 26A View Fig , 28A View Fig , 32A View Fig ); basomedially on T1–T3 hair bands of long, coarse, and dense hair ( Figs 25D View Fig , 27D View Fig , 31D View Fig )..... 19

18. Disc of T2 basally densely (i = 0.2 d) punctate, punctation covers at most basal half of tergum ( Fig. 53A View Fig ); propodeum apically carinate ( Fig. 52C View Fig ) .......................... S. sphecodoides ( Friese, 1912)

– Disc of T2 basally densely (i = 0.2 d) punctate, punctation covers at most basal three quarters of tergum ( Fig. 50A View Fig ); propodeum completely areolate ( Fig. 49C View Fig )........... S. semirufus Cockerell, 1932

19. Gonocoxites medial angular-shaped ( Figs 27B View Fig , 32B View Fig , 47B View Fig ); scutum completely densely (i <1 d) and regularly punctate ( Figs 26B View Fig , 31B View Fig , 46B View Fig ); disc of T2 apicomedially with at least 20 punctures ( Figs 28A View Fig , 32A View Fig , 47A View Fig )...................................................................................................................... 20

– Gonocoxites medially curved ( Figs 41B View Fig , 43B View Fig , 57B View Fig ); scutum not completely densely (i <1 d) punctate, punctation irregular ( Figs 40B View Fig , 42B View Fig , 56B View Fig ); disc of T2 apicomedially at most with 15 punctures ( Figs 41A View Fig , 45A View Fig , 57A View Fig )...................................................................................................... 22

20. Flagella of antennae about twice as long as compound eye ( Fig. 31A View Fig ) .......... S. longicornis sp. nov.

– Flagella of antennae about 1.4 × as long as compound eye ( Figs 28A View Fig , 46A View Fig ) ................................. 21

21. Disc of T2 basally densely (i = 0.2 d) punctate, punctation covers at most basal three quarters of tergum, getting sparser apically ( Fig. 47A View Fig ); posterior ocelli little behind posterior margin of compound eyes; distal margin of S7 with wide and deep emargination ( Fig. 47C View Fig )............................ ........................................................................................................ S. perpunctatus Cockerell, 1933

– Disc of T2 basally densely (i = 0.2 d) punctate, punctation covers at most basal half of tergum ( Fig. 28F View Fig ); posterior ocelli much behind posterior margin of compound eyes ( Fig. 28B View Fig ); S7 with slight emargination ( Fig. 29C View Fig )............................................................................ S. imparilis sp. nov.

22. T2 basally with dense punctation that covers more than basal half of tergum ( Fig. 43A View Fig ); discs of T2–T4 medially with irregular coarse punctures ( Fig. 42D View Fig ); propodeum completely and coarsely areolate ( Fig. 42C View Fig )..................................................................................... S. perpunctatulus sp. nov.

– T2 basally with dense punctation that covers basal third to half of tergum ( Figs 26A View Fig , 41A View Fig , 57A View Fig ); discs of T2–T4 medially with regular coarse punctures ( Figs 25D View Fig , 40D View Fig , 56D View Fig ); propodeum not completely areolate or with other type of sculpture ( Figs 25C View Fig , 40C View Fig , 56C View Fig )..................................... 23

23. Discs of T1–T3 finely punctate ( Fig. 40C View Fig ) ........................................................... S. obtusus sp. nov.

– Discs of T1–T3 coarsely punctate ( Figs 25D View Fig , 56D View Fig ) ........................................................................ 24

24. Propodeum laterally carinate, medially rugulose-areolate ( Fig. 56C View Fig )................. S. variabilis sp. nov.

– Basal third of propodeum coarsely areolate, apical part finely areolate or coriaceous ( Fig. 25C View Fig )...... .............................................................................................................. S. glaberrimus ( Friese, 1912)