Bergera kwangsiensis (C.C.Huang) F.J.Mou, 2023

Mou, Feng-Juan, Hu, Xiu, Ha, Bui Thu & Cuong, Nguyen Manh, 2023, Taxonomic revision of Bergera J. Koenig ex L. (Rutaceae) based on the molecular phylogeny and morphology, European Journal of Taxonomy 860, pp. 141-180 : 163-165

Bergera kwangsiensis (C.C.Huang) F.J.Mou , comb. nov.

urn:lsid:ipni.org:names:77314675-1

Fig. 8

Clausena kwangsiensis C.C.Huang, Acta Phytotaxonomica Sinica 8 (1): 90 ( Huang 1959). – Murraya kwangsiensis (C.C.Huang) C.C.Huang, Acta Phytotaxonomica Sinica 16 (2): 85 ( Huang 1978). – Type: CHINA • Guangxi, Baise, Baiguoshan; alt. 830 m; R.C. Ching 7667; holotype: PE[00022500] image!; isotype: NAS[NAS00404843] image!.

Etymology

The specific epithet is derived from the type locality, Guangxi in China.

Material examined

CHINA • Guangxib Province; 5 Sep. 1928; R. C. Ching 7667; NAS image ! • 1958; Nanning Institute of traditional Chinese Medicine 11744; GXMG ! • 1960; Nanning Institute of traditional Chinese Medicine 11758; GXMG ! • 1964; Y.X. Gan 68238; GXMG ! • 8 Jun. 1983; J. Y. Luo & J. X. Ling 76250; GXMI ! • 17 Dec. 1985; L. Ou & A. N. Mo 000748; GXMI ! • 25 Sep. 1990; D. Fang & L. Zeng 169; GXMI ! • Shanglin, Mushan; 9 Jun. 2014; J. B. Teng et al. 450125140609023 LY; GXMG ! • Long’an, Chengxiang; 19 Jan. 2013; Long’an General Expedition 450123130119033 LY; IBK ! • 9 Nov. 2013; Long’an General Expedition 450123130119033 LY; GXMG ! • 25 Jun. 2014; Long’an General Expedition 450123140625098 LY; IBK!, GXMG ! • Nanwei; 3 Dec. 2012; Long’an General Expedition 450123121203008 LY; IBK!, GXMG ! • Dujie; 6 Nov. 2013; Long’an General Expedition 450123130116012 LY; IBK!, GXMG ! • 27 Jun. 2014; Long’an General Expedition 450123140627072 LY; IBK!, GXMG ! • Pinshan, Longhushan; 24 Jul. 2013; Long’an General Expedition 450123130724004 LY; IBK ! • Longhushan; 13 Oct. 1981; Expedition 11321; GXMG ! • 11 May 2008; Guangxi Expedition of Chinese Plant Combination 0162; IBK ! • Xinguang; 12 Nov. 2011; J. C. Yang & Y. B. Liao LH 1316; IBK ! • Yali; 22 Nov. 2011; J. C. Yang & Y. B. Liao LH 1765; IBK ! • Wuming; 23 Jun. 1955; Guangxi Team 528; PE image ! • 16 Dec. 1991; B. S. Huang 22767; GXMI ! • Tengxiang ; Jun. 1970; S. Q. He s.n.; IBK ! • Taiping ; 29 Jan. 1975; J. J. Wang 6578; GXMI ! • 21 Jul. 1985; Q. Li 166; SYS ! • 14 Nov. 1985; Q. Li 173; SYS ! • Chengxiang; 12 Decembr 2013; J. B. Teng et al. 450122131212004 LY; GXMG ! • Ningwu ; 14 Jul. 2010; K. J. Yan 22707; GXMI ! • 19 Nov. 2014; J. B. Teng et al. 450122141119007 LY; GXMG ! • Yongning ; 26 Jul. 1958; S. Q. Zhong A62018 ; KUN!, IBK ! • 16 Sep. 1964; J. Q. Ge & Y. Wang 17638; GXMI ! • Dingtun; 20 Jul. 1985; Q. Li 167; SYS ! • Chongzuo, Heping; 13 Nov. 1997; D. R. Liang & B. S. Huang 02570; GXMI ! • Ningming; 14 Dec. 1977; G.D. Guo 15489; HITBC ! • Nongrui; 29 Apr. 1980; Nonggang Comprehensive Expedition 11881; IBK ! • 26 Jul. 2011; Y. S. Huang & Y. B. Liao NM1369 ; IBK ! • 30 May 2013; B. Y. Huang et al. 451422130530024 LY; GXMG ! • Tuolong; 8 Dec. 2012; H. Z. Lü et al. 451422121208075 LY; GXMG ! • Tingliang; 2 Nov. 2013; H. Z. Lü et al. 451422131102031 LY; GXMG ! • Chengzhong; 26 Dec. 2012; B. Y. Huang et al. 451422141226027 LY; GXMG ! • 4 Nov. 2013; H. Z. Lü et al. 451422131104016 LY; GXMG ! • Longzhou, Jinlong ; 3 Jul. 1960; C. J. Liang 21533; GXMI ! • Shangjin; 31 Oct. 1979; Nonggang Comprehensive Expedition 20565; GXMI ! • 17 Aug. 1989; K. J. Yan & W. Y. Rao 76364; GXMI ! • Nonggang, Nonghu; 24 Sep. 1979; Nonggang Comprehensive Expedition 10310; IBK ! • Shida; 5 Oct. 1979; Nonggang Comprehensive Expedition 10790; IBK ! • Pingxiang, Xiashi; 21 Dec. 1959; X. F. Deng 10670; IBK ! • Xiangshui; 8 Jun. 1979; Investigation team of Longzhou Medical Institute 0024; GXMI ! • 14 Jun. 1979; Investigation team of Longzhou Medical Institute 0264; GXMI ! • 30 Dec. 1986; Y. P. Tao 80172; GXMI ! • Daxin, Taiping, Anmin ; 21 Jun. 1977; Y. Z. Zhang Da-127; GXMI ! • Longlin; 15 Apr. 1979; S. Q. Lin & M. F. Qin 2132; IBK ! • Youjiang, Banshui; 18 Jul. 2014; Youjiang General Expedition 451002140718024 LY; GXMG ! • Jinxi, Kuiwei, Dezhou ; 11 Oct. 2013; X. Y. Huang et al. 451025131011014 LY; GXMG ! • Napo Dingye ; 20 Jun. 2010; Y. F. Huang & J. Huang HYF0165; GXMG ! • Yunnan Province, Funing, Bo’ai ; 21 Oct. 1958; H. T. Tsai 8813; KUN ! • Guichao ; 4 Mar. 2007; F. J. Mou 75, 77; IBSC ! • Guichao ; 20 Jul. 2007; F. J. Mou 140; IBSC ! • 19 Sep. 2016; F. J. Mou 528; SWFC ! • Wenshan, Xishan ; 14 Oct. 1958; H. T. Tsai 8444; KUN ! • Mengla, Menglun , cultivated; 3 Feb. 2007; F. J. Mou 41, 42; IBSC !.

Description

Shrub, 1–2 m tall. Leaves are 3–11-foliolate, 3–10 × 2.5–6.5 cm; rachises and petiolules covered with soft hairs; leaflet blades alternate, ovate-oblong, coriaceous, dispersively oil-glandular, dark green and shining adaxially and yellowish green abaxially, lower surface covered densely with white soft hairs abaxially, apex obtuse to rounded, margin crenulate; petiolules 2–3 mm. Inflorescences are terminal. Flowers are 5-merous, ellipsoid in bud; sepals broadly ovate, ca 1 mm; petals ca 4 mm, dispersing oil glands. Stamens are 10; filaments flat and widening down; filaments and anthers ciliate. Ovaries are oblong, as long as styles; stigmas not prominent. Fruits are globose, ca 1 cm in diam., and red when ripe.

Phenology

Flowering between June and July, while fruiting from October to November.

Distribution and habitat

China: Guangxi (Longzhou, Ningming, Wuming, Yongning, Fushui, and Baise), Yunnan (Wenshan and Funing); Vietnam: Quỳ Chau. Growing in the shrub or forestry of limestone areas, alt. 100–800 (1400) m.

Remarks

The leaves and fruits of this species have a strong, pleasant scent.

Figures

Fig. 8. Bergera kwangsiensis (C.C.Huang) F.J.Mou comb. nov. A. Plant. B. Inflorescence. C. Flowers and young fruits. D–E. Infructescence. F. Pistil. G. Stamens. H. Ovary crossection. Photos taken by Feng-Juan Mou in China.