Forcipomyia (Metaforcipomyia) macroseta, Spinelli, Gustavo R., Marino, Pablo I. & Borkent, Art, 2012

Spinelli, Gustavo R., Marino, Pablo I. & Borkent, Art, 2012, A revision of Biting Midges of the Subgenera Forcipomyia (Metaforcipomyia) and F. (Saliohelea) from Costa Rica (Diptera: Ceratopogonidae), Zootaxa 3419, pp. 1-52 : 20-21

publication ID	https://doi.org/ 10.5281/zenodo.215031
DOI	https://doi.org/10.5281/zenodo.6166124
persistent identifier	https://treatment.plazi.org/id/03A7878C-E365-FFD2-53D2-BFBA8032FF25
treatment provided by	Plazi
scientific name	Forcipomyia (Metaforcipomyia) macroseta
status	sp. nov.

Treatment

Forcipomyia (Metaforcipomyia) macroseta View in CoL n. sp.

( Figs. 57–61 View FIGURES 57 – 61 , 113 View FIGURES 112 – 115 , 141 View FIGURES 140 – 141 )

Diagnosis. Male: only species of Neotropical Forcipomyia (Metaforcipomyia) with a hind tarsal ratio of 2.10, flagellomere 10 two X length of flagellomere 11, tarsomere 2 of foreleg with thick, reinforced setae, and parameres present. Female: unknown.

Description. Male. Head. Brown. Eyes abutting medially for length of four ommatidia. Antenna ( Fig. 57 View FIGURES 57 – 61 ) light brown except flagellomere 1 brown, with plume setae well developed, flagellomeres 2–4 elongate, flagellomere 10 two X longer than flagellomere 11, flagellomeres 10–13 elongate; flagellomere 13 with apical nipple, slightly constricted basally; AR 0.91. Palpus ( Fig. 58 View FIGURES 57 – 61 ) brown; segment 3 slightly swollen proximally (or on 2/3 proximal), with shallow sensory pit opening at midlength; PR 2.20. Labrum tapering apically.

Thorax. Uniformly dark brown; scutellum with 8 strong setae, and several minor ones. Legs brown, apex of hind tibia with 3–4 spines; tarsomeres with pectinate scales; tarsomere 1 of foreleg with row of dozen spine-like bristles, tarsomere 2 of foreleg with 1–2 thick reinforced setae, tarsomere 3 of foreleg with one setae weaker that latter ones, tarsomere 1 of hind leg with two rows of thick setae, tarsomeres 2–5 slightly paler; foreleg TR 2.75, midleg TR 2.00, hind leg TR 2.10; claws curved, moderately stout. Wing ( Figs. 59 View FIGURES 57 – 61 , 113 View FIGURES 112 – 115 ) with pattern of pigmented membrane, with dark patch on its anterior margin and at the apices of M2, CuA1 and CuA2; M2 only visible at wing margin; first radial cell obliterated; second radial cell well developed, triangular; fork of cubitus situated just at level of apex of costa; wing length 0.90 mm; breadth 0.30 mm; CR 0.42. Halter whitish.

Abdomen. Tergites brown. Genitalia ( Fig. 60 View FIGURES 57 – 61 ): Dark brown except gonostyli pale. Tergite 9 short, only extending to 1/4 length of gonocoxite; posterior margin broadly rounded; cercus lobe-like, produced beyond midlength of gonocoxite; sternite 9 broad, nearly straight. Gonocoxite stout, two X longer than greatest breadth; gonostylus 0.85 length of gonocoxite, almost straight, abruptly tapering to sharply pointed tip. Parameres represented by a slender, arrow-shaped, posteromesally directed projection, gonocoxal apodemes nearly straight, narrow. Aedeagus ( Fig. 61 View FIGURES 57 – 61 ) triangular, slightly tapering to pointed tip (triangular apex folded in the available specimen), arrowhead- shaped, sharply blunt tip; basal arch extending to 1/4 of total length; lateral arms extending laterally.

Female. Unknown.

Distribution. This species is known only from the type locality in Costa Rica, at 930 meters ( Fig. 141 View FIGURES 140 – 141 ).

Taxonomic discussion. This species shares with F. longiflagellata the presence of thick, reinforced setae on tarsomere 2 of the foreleg and the similarly shaped parameres. However, the wing of the latter species is plain and the aedeagus tapers slightly to a broad, truncate tip. It is also similar to F. grandiseta which shares the presence of thick, reinforced setae on tarsomere 2 of the foreleg, but differs from this species by the stouter aedeagus, which tapers to a broad posterior margin and bears two well developed apical lobes. The Nearctic species F. fehrerorum Grogan & Sigrist is nearly identical to F. macroseta , but the aedagus have an underlying hyaline pointed structure, the basal arch is lower and the apex is bifid.

Type material. Holotype male on microscope slide, labeled “ HOLOTYPE Forcipomyia (Metaforcipomyia) macroseta Spinelli, Marino and Borkent , Costa Rica, Limón prov., Talamanca, La Amistad NP entre Laguna Doroteri y Dabagri, 930 m, 23/ 27-VII-2007, A. Solis – M. Moraga, Malaise tp. LS 396680_543377 #92378” ( INBC).

Derivation of specific epithet. The name macroseta – macro (large), seta refers to the stout setae on the foreleg tarsomere 2 of the male of this species.