Tomosvaryella freidbergi De Meyer, 1995

Motamedinia, Behnam, Skevington, Jeffrey H. & Kelso, Scott, 2021, Revision of Tomosvaryella Aczél (Diptera: Pipunculidae) in the Middle East, with description of 19 new species, Zootaxa 5002 (1), pp. 1-103 : 30

Tomosvaryella freidbergi De Meyer, 1995

Figs 14A‒E , 54

Diagnosis: This species can be recognized by the slender shape of surstyli in dorsal view ( Fig. 14A ); both surstyli straight in basal half, gently bent in apical half in lateral view ( Fig. 14D‒E ); epandrium longer than wide ( Fig. 14A ); gonopods equal in height ( Fig. 14B ). Based on the shape of genitalia, this species is part of the kuthyi species group ( Tomosvaryella emaratensis sp. nov., T. cyprusensis sp. nov., T. hispanica , T. israelensis , T. kuthyi and T. parakuthyi ). It differs from these species by the shape of surstyli in lateral view, slender, slightly straight in basal half and being bent in apical half ( De Meyer, 1995: fig. 13a–c; Földvári & De Meyer, 1999: fig. 10A–D). See diagnosis of T. emaratensis sp. nov. and T. cyprusensis sp. nov. for more details.

Specimens examined: HOLOTYPE: ISRAEL: Mt. Hermon , 33°25’N, 35°51’E, 2000 m a.s.l., 1.vii.1986, A. Freidberg, JSS52375 (1♂, TAU) ; PARATYPE: ISRAEL: Meron , 32°59’N, 35°26’E, 11.iv.1974, F. Nachbar, JSS52376 (1♂, TAU) . Other specimens examined: CYPRUS: Kyrenia, 35°21’N, 33°09’E, 16‒30.vii.2017, O. Ozden, Malaise trap, JSS52356 (1♂, CNC) ; IRAN: North Khorasan: Biar , 37°53’N, 57°35’E, 1008 m a.s.l., 10‒ 24.vi.2016, B. Motamedinia, Malaise trap; JSS52018 (1♂, CNC) ; Chenaran village , 37°27’N, 57°36’E, 1300 m a.s.l., 12.vi.2016, B. Motamedinia, sweep net, JSS51963 (1♂, CNC) ; South Khorasan: Ghiok , 32°47’N, 59°07’E, 1917 m a.s.l., 9.vi.2016, B. Motamedinia, sweep net, JSS52000 (1♂, CNC) ; Kermanshah: Dodan, 35°00’N, 46°12’E, 1011 m a.s.l., Garden , 20.vi.2016, M. Zardouei, Malaise trap, JSS52177 (1♂, CNC) ; ISRAEL: Har Hermon , 33°18’N, 35°46’E, 2000 m a.s.l., 3.viii.1995, A. Freidberg, JSS52076 (1♂, TAU) .

Distribution: Czech Republic Egypt, France, Greece, Hungary, Iran, Israel, Italy, Kyrgyz Republic, Portugal, Spain, Syria, Turkey ( Skevington 2020) ( Fig. 54 ).

Notes: DNA barcodes of T. freidbergi are identical to those of T. kuthyi (0.0‒1.5% pairwise divergence). The terminalia of these species differ a little by the shape of left surstylus, so this is likely a case of recently-diverged species or ancestral hybridisation. It is possible that these are a single species with polymorphic genitalia, so future genetic work is warranted.

Figures

FIGURE 14. Male genitalia of Tomosvaryella freidbergi, holotype (JSS52376, TAU) A) dorsal view, B) ventral view, C) ejaculatory apodeme, D, E) lateral view.

FIGURE 54. Distribution of Tomosvaryella freidbergi, T. saudiensis sp. nov., T. subtransvaalensis sp. nov. in the Middle East.