Haasea flavescens ( Latzel, 1884 )

Antić, Dragan & Akkari, Nesrine, 2020, Haasea Verhoeff, 1895 - a genus of tumultuous history and chaotic recordsredefinition, revision of taxonomy and geographic distributions, with descriptions of two new species from Austria and Serbia (Diplopoda, Chordeumatida, Haaseidae), Zootaxa 4798 (1), pp. 1-77 : 18-25

publication ID	https://doi.org/ 10.11646/zootaxa.4798.1.1
publication LSID	lsid:zoobank.org:pub:4830ED2F-DB55-468E-97D4-2A278CBA6CE0
persistent identifier	https://treatment.plazi.org/id/03A90C3B-152A-FF90-FF24-72B4FBAEFCA4
treatment provided by	Plazi
scientific name	Haasea flavescens ( Latzel, 1884 )
status

Treatment

Haasea flavescens ( Latzel, 1884) View in CoL

Figs 1 View FIGURE 1 C–D, 2, 3D, H, J, 8–10, 38, 39

Craspedosoma flavescens Latzel, 1884 :

Craspedosoma flavescens helveticum Verhoeff, 1894 View in CoL — Schubart (1934: 87)

Orobainosma pinivagum Verhoeff, 1901 — Spelda (2005: 121)

Orobainosoma flavescens setosum Rothenbühler, 1900 View in CoL — Verhoeff (1910b: 52)

Orobainosoma flavescens vornatscheri Verhoeff, 1935 View in CoL syn. nov.

Diagnosis. Differs from the other congeners, except H. hungarica , by the presence of the distal notch of the posterior lamella. From H. hungarica , H. flavescens differs by 1) the number of body segments (mainly 30 in H. flave- scens vs. 28 in H. hungarica ), 2) the shape and size of the posterior lamella (smaller without proximal protrusion in H. flavescens vs. larger with proximal protrusion in H. hungarica ) and 3) the shape of mesal process (distal margin densely denticulated with an isolated posterior branch with a small tooth in H. flavescens vs. distal margin with several teeth, and with fork-like end in H. hungarica ).

Type material examined: Orobainosma pinivagum Verhoeff, 1901 (marked as a syntypes): ZMB12737: legpairs 1–7, ZMB12738: anterior gonopods, posterior gonopods, leg-pair 7, leg-pairs 10–11, ZMB12739: leg-pair 10, anterior gonopods, posterior gonopods, leg-pair?12; Orobainosoma flavescens vornatscheri Verhoeff, 1935 (marked as syntypes), 2 juveniles, Türkenloch, 18.04.1935, Verhoeff det., Vornatscher leg. ( NHMW 9500).

Other material examined: AUSTRIA-HUNGARY:several specimens,1919, R.Latzel leg.,don.( NHMW 9546); 1 fragment (anterior gonopod), same area, 1919, R. Latzel leg., don. ( NHMW9549 View Materials ) ; AUSTRIA: Burgenland: Eisenstadt-Umgebung : 5 juveniles, Hornstein E mountain valley, ca 320 m, 14.08.1998, J. Gruber leg. ( NHMW9758 View Materials ) ; 3 juveniles, Loretto 1.4 km S- wine trail, ca 320 m, 14.08.1998, J. Gruber leg. ( NHMW9743 View Materials ) ; Lower Austria: Amstetten : 2 juveniles, Hilm-Ybbs flood plains, ca 305 m, 25.09.2013, J. Gruber leg. ( NHMW9741 View Materials ) ; 1 juvenile, Kreuzbergweg , ca 300 m, 14.09.1997, J. Gruber leg. ( NHMW9737 View Materials ) ; 1 juvenile, same locality, 21.08.2010, J. Gruber leg. ( NHMW9763 View Materials ) ; 1 juvenile, Zauchbach (Kröllendorf E), ca 325 m, 18.09.2012, J. Gruber leg. ( NHMW9773 View Materials ) ; Baden : 1 juvenile, “Eugen- Villa” N- Schwechat river bank, ca 245 m, 02.06.1990, J. Gruber leg. ( NHMW9755 View Materials ) ; 1 ♂, C. Attems det., Alland , 09.08.1936, J. Vornatscher leg. ( NHMW9505 View Materials ) ; 2 juveniles, Lerchenfeld N slope ( Furth Triesting ), ca 390 m, 24.08.1996, J. Gruber leg. ( NHMW9742 View Materials ) ; 1 juvenile, Steinwandgraben (Ebeltaler Bach S), ca 430 m, 24.08.1996, J. Gruber leg. ( NHMW9760 View Materials ) ; Bruck an der Leitha : 3 juveniles, Hainburger Berge- N to “Rotes Kreuz”, ca 250 m, 12.08.1989, J. Gruber leg. ( NHMW9736 View Materials ) ; 1 ♂, C. Attems det., Hundsheimer Berg ( Bad Deutsch Altenburg ), 14.10.1924,( NHMW9486 View Materials ) ; 1♀, C.Attems det., Mannersdorf—Kaisereiche , 14.10.1924, ( NHMW9485 View Materials ) ; Kirchberg am Wechsel : 4 juveniles, 1 ♂, 1 ♀, C. Attems det., Hermannshöhle , 04.06.1905, ( NHMW9478 View Materials ) ; several specimens, C. Attems det., same locality, 18.04.1935, J. Vornatscher leg. ( NHMW9503 View Materials ) ; several specimens, K.W. Verhoeff det., same locality, 13.02.1938, J. Vornatscher leg. ( NHMW9502 View Materials ) ; Klosterneuburg : 1 fragment , 3 juveniles, Gugging river valey “Lourdesgrotte” 500 m N, ca 275 m, 03.07.1983, J. Gruber leg. ( NHMW9746 View Materials ) ; Korneuburg : 3 juveniles, Mühlberg W ( Hausleiten N), ca 190 m, 11.07.1998, J. Gruber leg. ( NHMW9748 View Materials ) ; Krems-Land : 2 juveniles, Bacharnsdorf S, ca 240 m, 17.07.2004, J. Gruber leg. ( NHMW9738 View Materials ) ; 1 juvenile, same locality, 26.09.2003, J. Gruber leg. ( NHMW9754 View Materials ) ; 2 ♀♀, C. Attems det., Spitz beechforest, 19.10.1938, ( NHMW9488 View Materials ) ; Liezen : 1 juvenile, Königstetten S- creek near Kemmelbach , ca 250 m, 11.09.1999, J. Gruber leg. ( NHMW9739 View Materials ) ; Lilienfeld : 1 juvenile, Dickenau gorge (Türnitz NE), ca 440 m, 19.08.1990, J. Gruber leg. ( NHMW9740 View Materials ) ; 1 juvenile, Fliedersbach-Au ( Hainfeld ), ca 440 m, 28.07.1994, J. Gruber leg. ( NHMW9759 View Materials ) ; 1 juvenile, Sankt Veit an der Gölsen- valey NW 12 km, ca 285 m, 31.07.2009, J. Gruber leg. ( NHMW9762 View Materials ) ; 8 specimens, St. Aegyd am Neuwalde- slope S, ca 570 m, 08.07.2010, J. Gruber leg. ( NHMW9765 View Materials ) ; 3 juveniles, same area-Seebachstr., ca 590 m, 08.07.2010, J. Gruber leg. ( NHMW9772 View Materials ) ; 1 ♂, C. Attems det., Türkenloch bei Hainfeld , 08.12.1936, J. Vornatscher leg. ( NHMW9506 View Materials ) ; 1 juvenile, valley in W slope (cemetary S), ca 430 m, 24.08.2009, J. Gruber leg. ( NHMW9770 View Materials ) ; 5 juveniles, same locality, 18.08.2010, J. Gruber leg. ( NHMW9771 View Materials ) ; 17 ♂♂, 30 ♀♀, same locality, 12.10.2010, J. Gruber leg. ( NHMW9733 View Materials ) ; Melk : 1 juvenile, Hubertendorf- Graben N, ca 250 m, 12.08.1998, J. Gruber leg. ( NHMW9751 View Materials ) ; 1 juvenile, Mank-Au ( Hörsdorf S), ca 285 m, 13.08.2007, J. Gruber leg. ( NHMW9745 View Materials ) ; 2 juveniles, Pühra- creek ( St. Leonhard am Forst ), ca 260 m, 31.08.2009, J. Gruber leg. ( NHMW9761 View Materials ) ; 2 juveniles, Weitenegg N to ruins- at N slope, ca 220 m, 29.07.2001, J. Gruber leg. ( NHMW9735 View Materials ) ; 16 juveniles, Ysperklamm- sidecreek S, ca 630 m, 30.08.1990, J. Gruber leg. ( NHMW9757 View Materials ) ; Mödling : 1 juvenile, “ Geschriebene Buche ” W 500 m (Gruberau S), ca 470 m, 01.09.1991, J. Gruber leg. ( NHMW9753 View Materials ) ; 3 ♂♂, 4 ♀♀, C. Attems det., Anninger , 01.11.1938, ( NHMW9487 View Materials ) ; 1 specimens, Kaltenleutgeben , 28.07.1879, Feiller leg. ( NHMW9524 View Materials ) ; Neunkirchen : 1 juvenile, Grimmenstein NE Raintal-side valley , ca 500 m, 12.08.2000, J. Gruber leg. ( NHMW9756 View Materials ) ; Sankt Pölten: 5 juveniles, city forest, ca 290 m, 15.05.1999, J. Gruber leg. ( NHMW9747 View Materials ) ; 1 juvenile, Unterradlberg- Hainerstrasse , ca 250 m, 16.09.2005, J. Gruber leg. ( NHMW9768 View Materials ) ; Sankt Pölten-Land: 2 ♂♂, C. Attems det., “ In der Au am Krak- ing”, 10.1914, ( NHMW9480 View Materials ) ; 1 juvenile, “ Oberer Dambach ” (Ollersbach S), ca 300 m, 08.07.2007, J. Gruber leg. ( NHMW9749 View Materials ) ; 30 ♂♂, 30 ♀♀, C. Attems det., Kirchberg an der Pielach , 02.10.1924, ( NHMW9484 View Materials ) ; 1 juvenile, Perschenegg- Perschling river , ca 370 m, 20.08.1990, J. Gruber leg. ( NHMW9744 View Materials ) ; 3 juveniles, same locality, 20.08.1990, J. Gruber leg. ( NHMW9734 View Materials ) ; 9 juveniles, Pielach-Au ( Obergrafendorf S), ca 280 m, 23.08.1990, J. Gruber leg. ( NHMW9750 View Materials ) ; 1 juvenile, Plosdorf-flood plain forest Michelbach , ca 240 m, 20.08.1988, J. Gruber leg. ( NHMW9767 View Materials ) ; 2 ♂♂, 3 ♀♀, C. Attems det., Rekawinkel , 27.10.1893, ( NHMW9477 View Materials ) ; 1 juvenile, Schildberg (Böheimkirchen N), ca 270 m, 16.08.1998, J. Gruber leg. ( NHMW9764 View Materials ) ; 1 juvenile, Schildberg (Weischling SW), 16.08.1998, J. Gruber leg. ( NHMW9766 View Materials ) ; 1 juvenile, Tobelgraben NE ( Seelackenberg ), ca 250 m, 12.06.1998, J. Gruber leg. ( NHMW9774 View Materials ) ; 2 juveniles, Traisen-Au between Göblasbruck & Kreisbach, 15.08.1989, J. Gruber leg. ( NHMW9752 View Materials ) ; Scheibbs : 7 juveniles, C. Attems det., Gaisenberg bei Puchenstuben , 1947, H. Strouhal leg. ( NHMW9507 View Materials ) ; 1 ♂, Gaming , 1883, Feiller leg. ( NHMW9517 View Materials ) ; 1 ♀, C. Attems det., Puchenstuben , 01.10.1924, ( NHMW9483 View Materials ) ; 8 ♂♂, 5 ♀♀, C. Attems det., Seehof to Rehberg ( Lunz am See ), 25.09.1924 - 27.09.1924, ( NHMW9481 View Materials ) ; 2 ♀♀, C.Attems det., Trübenbach , 30.09.1924, ( NHMW9482 View Materials ) ; 2 ♂♂, C. Attems det., Wilhelmshöh- le, 1939, Troll leg. ( NHMW9504 View Materials ) ; Tulln : 1 specimens, Kierling , 23.10.1884, Feiller leg. ( NHMW9522 View Materials ) ; 1 juvenile, Schloss Hadersfeld W 1 km, ca 350 m, 01.09.1985, J. Gruber leg. ( NHMW9769 View Materials ) ; Wiener Neustadt-Land: 1 ♂, C. Attems det., Dürre Wand, 10.1906, ( NHMW9479 View Materials ) ; 2 ♂♂, 1 ♀, C.Attems det., Krumbach N, Troll leg. ( NHMW9501 View Materials ) ; Styria: Bruck-Mürzzuschlag : 1 ♀, C. Attems det., Hochschwab Bürgeralpe, ( NHMW9490 View Materials ) ; Hartberg-Fürstenfeld : 1 juvenile, Tauchen S, 10.09.1947, H. Strouhal leg. ( NHMW9514 View Materials ) ; 4 fragments, 2 juveniles, 8 ♂♂, 2 ♀♀, C. At- tems det., Vorau , 14.10.1909, ( NHMW9492 View Materials ) ; Liezen : 2 ♂♂, C. Attems det., Ardning bei Admont , 28.10.1941, H. Franz leg. ( NHMW9511 View Materials ) ; 1 ♂, C. Attems det., Dürrnschöberl , H. Franz leg. ( NHMW9512 View Materials ) ; 2 ♂♂, 1 ♀, C. Attems det., Hall near Admont , 10.11.1941, H. Franz leg. ( NHMW9510 View Materials ) ; 2 ♂♂, C. Attems det., Kemmatgraben near Ad- mont- shieling (Hochalm), 1940, H. Franz leg. ( NHMW9509 View Materials ) ; Upper Austria: Steyr-Land : 1 ♂, C. Attems det., Almkogelgipfel , 1943, H. Franz leg. ( NHMW9508 View Materials ) ; Vienna: 1 ♂, C. Attems det., Orobainosoma flavescens J. Gruber rev., A217, H. Franz leg. ( NHMW9466 View Materials ) ; 2 specimens, Donauauen left riverside, 14.11.1881, Feiller leg. ( NHMW9523 View Materials ) ; 3 ♂♂, 6 ♀♀, C. Attems det., Lobau , 06.11.1893, ( NHMW9496 View Materials ) ; Döbling : 1 ♂, Bellevuestreet & Kaasgrabengasse, ca 330 m, 06.02.2000, J. Gruber leg. ( NHMW9710 View Materials ) ; 7 juveniles, Cobenzl ( Latisberg ) E slope, ca 400 m, 09.08.1980, J. Gruber leg. ( NHMW9698 View Materials ) ; 1 ♀, Cobenzl ( Latisberg ) NE slope, ca 375 m, 19.11.1994, J. Gruber leg. ( NHMW9699 View Materials ) ; 1 ♀, Gspöttgraben , 320 m, 15.03.1981, J. Gruber leg. ( NHMW9687 View Materials ) ; 1 juvenile, same locality, 24.05.1981, J. Gruber leg. ( NHMW9688 View Materials ) ; 1 ♂, 1 ♀, Hackenberg N slope, ca 290 m, 06.01.1982, J. Gruber leg. ( NHMW9708 View Materials ) ; 1 ♂, Kahlenberg E slope, ca 280-300 m, 04.01.1981, J. Gruber leg. ( NHMW9693 View Materials ) ; 1 ♂, same area, ca 410 m, 04.01.1980, J. Gruber leg. ( NHMW9692 View Materials ) ; 2 juveniles, Krapfenwaldl , ca 360 m, 12.09.1985, J. Gruber leg. ( NHMW9689 View Materials ) ; 4 ♂♂, 6 ♀♀, same locality, 20.10.1985, J. Gruber leg. ( NHMW9690 View Materials ) ; 2 ♂♂, 18 ♀♀, same locality, 07.12.1985, J. Gruber leg. ( NHMW9691 View Materials ) ; 1 ♀, Kronesgasse , ca 220 m, 22.12.1985, J. Gruber leg. ( NHMW9683 View Materials ) ; 1 ♀, Leopoldsberg N slope, ca 380 m, 26.10.1985, J. Gruber leg. ( NHMW9709 View Materials ) ; 3 juveniles, 6 ♂♂, 8 ♀♀, C. Attems det., Leopoldsberg S upper slope, ca 390m, 11.10.1980, J. Gruber leg. ( NHMW2889 View Materials ) ; 1 ♂, 1 ♀, N slope (N to Hofzeile ), ca 180 m, 04.01.1981, J. Gruber leg. ( NHMW9696 View Materials ) ; 1 ♂, 1 ♀, same locality, 11.12.1988, J. Gruber leg. ( NHMW9664 View Materials ) ; 2 ♂♂, same locality, 01.04.1990, J. Gruber leg. ( NHMW9665 View Materials ) ; 2 juveniles, same locality, 04.09.1999, J. Gruber leg. ( NHMW9697 View Materials ) ; 1 ♂, NW Obersteinergasse , 190 m, 08.02.1981, J. Gruber leg. ( NHMW9666 View Materials ) ; 6 ♂♂, 7 ♀♀, Obere Wildgrube NE slope ( Schwabenwiese ), ca 400 m, 28.12.1980, J. Gruber leg. ( NHMW9701 View Materials ) ; 1 ♀, same locality, 31.12.1994, J. Gruber leg. ( NHMW9704 View Materials ) ; 2 juveniles, Obere Wildgrube NE slope, ca 300 m, 15.09.1987, J. Gruber leg. ( NHMW9702 View Materials ) ; 5 juveniles, same area, ca 400 m, 20.09.1980, J. Gruber leg. ( NHMW9700 View Materials ) ; 1 ♀, same area, ca 425 m, 14.10.1989, J. Gruber leg. ( NHMW9703 View Materials ) ; 4 ♂♂, 6 ♀♀, same local- ity, 04.12.1999, J. Gruber leg. ( NHMW9705 View Materials ) ; 1 ♂, 2 ♀♀, Pointenbach lower valley ( Reisenbergweg ), ca 260 m, 24.02.1980, J. Gruber leg. ( NHMW9667 View Materials ) ; 1 juvenile, 1 ♂, same locality, 13.04.1980, J. Gruber leg. ( NHMW9668 View Materials ) ; 40 juveniles, same locality, 20.07.1980, J. Gruber leg. ( NHMW9670 View Materials ) ; 34 juveniles, same locality, 20.07.1980, J. Gruber leg. ( NHMW9669 View Materials ) ; 24 ♂♂, 3 ♀♀, same locality, 02.11.1980, J. Gruber leg. ( NHMW9671 View Materials ) ; 1 ♂, same local- ity, 31.12.1980, J. Gruber leg. ( NHMW9672 View Materials ) ; 18 ♂♂, 6 ♀♀, same locality, 17.10.1981, J. Gruber leg. ( NHMW9673 View Materials ) ; 13 juveniles, same locality, 19.06.1982, J. Gruber leg. ( NHMW9674 View Materials ) ; 6 ♂♂, 7 ♀♀, same locality, 14.11.1982, J. Gruber leg. ( NHMW9675 View Materials ) ; 3 juveniles, same locality, 25.08.1985, J. Gruber leg. ( NHMW9677 View Materials ) ; 10 ♂♂, same lo- cality, 26.10.1984, J. Gruber leg.( NHMW9676 View Materials ) ; 7 ♂♂, 4♀♀, same locality, 01.01.1988, J. Gruber leg.( NHMW9679 View Materials ) ; 7 ♂♂, 6 ♀♀, same locality, 15.12.1985, J. Gruber leg. ( NHMW9678 View Materials ) ; 1 juvenile, same locality, 05.06.1988, J. Gru- ber leg. ( NHMW9680 View Materials ) ; 3 ♂♂, 7 ♀♀, same locality, 17.12.1989, J. Gruber leg. ( NHMW9681 View Materials ) ; 3 ♂♂, 4 ♀♀, same locality, 28.12.1997, J. Gruber leg. ( NHMW9682 View Materials ) ; 4 ♂♂, 4 ♀♀, Pointenbach upper valley, ca 340 m, 19.10.1980, J. Gruber leg. ( NHMW9694 View Materials ) ; 1 ♀, same locality, 31.12.1980, J. Gruber leg. ( NHMW9695 View Materials ) ; 1 ♂, Rothschildgärten E slope, ca 180 m, 16.11.1980, J. Gruber leg. ( NHMW9663 View Materials ) ; 1 ♀, same area, ca 190 m, 28.09.1977 - 29.10.1977, J. Gruber leg. ( NHMW9684 View Materials ) ; 1 ♀, same locality, 29.10.1977, J. Gruber leg. ( NHMW9685 View Materials ) ; 1 ♀, same locality, 08.12.1977 - 05.03.1978, J. Gruber leg. ( NHMW9686 View Materials ) ; 3 juveniles, Steinbergbachtal ( Latisberg ), ca 380 m, 28.08.1982, J. Gruber leg. ( NHMW9706 View Materials ) ; 1 ♂, same locality, 22.10.1983, J. Gruber leg. ( NHMW9707 View Materials ) ; Hernals : 1 specimen, Neuwaldegg , 11.11.1881, Feiller leg. ( NHMW9526 View Materials ) ; 1 specimen, Neuwaldegg , 26.07.1882, Feiller leg. ( NHMW9527 View Materials ) ; 1 ♂, Neuwaldegg , 04.10.1883, Feiller leg. ( NHMW9525 View Materials ) ; 1 specimen, same locality, 04.10.1883, Feiller leg. ( NHMW9529 View Materials ) ; 1 ♀, Neuwaldegg- park burial chapel, 02.04.1883, Feiller leg. ( NHMW9528 View Materials ) ; Hietzing : 1 ♂, 1 ♀, Gelberberg E-slope (S Weidlingau), ca 300 m, 29.12.1979, J. Gruber leg. ( NHMW9650 View Materials ) ; 2 specimens, Royal garden Schönbrunn , “Römische Ruine”, 1885, Feiller leg. ( NHMW9545 View Materials ) ; 1 ♂, same area, ca 220 m, 16.12.1979, J. Gruber leg. ( NHMW9651 View Materials ) ; 5 ♂♂, 2 ♀♀, same locality, 14.10.1980, J. Gruber leg. ( NHMW9652 View Materials ) ; Leopoldstadt : 1 juvenile, 69 ♂♂, 48 ♀♀, Heustadelwasser N floodplain forest, ca 155m, 23.10.1980, J. Gruber leg. ( NHMW2886 View Materials ) ; 5 ♂♂, 2 ♀♀, Krebsenwasser S floodplain forest, ca 155 m, 30.10.1980, J. Gruber leg. ( NHMW9713 View Materials ) ; 2 juveniles, 2 ♂♂, 4 ♀♀, Lusthaus N floodplain forest, ca 155m, 07.10.1980, J. Gruber leg. ( NHMW9711 View Materials ) ; 13 ♂♂, 6 ♀♀, Lusthaus W floodplain forest, ca 155m, 14.12.1980, J. Gruber leg. ( NHMW2887 View Materials ) ; 1 ♂, Mauthnerwasser S floodplain forest, ca 155 m, 10.10.1980, J. Gruber leg. ( NHMW9712 View Materials ) ; 2 ♂♂, 1 ♀, same locality, 25.10.2000, J. Gruber leg. ( NHMW9715 View Materials ) ; 1 ♀, same locality, 25.10.2000, J. Gruber leg. ( NHMW9714 View Materials ) ; 1 ♀, Prater , Feiller leg. ( NHMW9530 View Materials ) ; 1 specimen, same locality, 1881, Feiller leg. ( NHMW9531 View Materials ) ; 2 specimens, same locality, 25.10.1881, Feiller leg. ( NHMW9532 View Materials ) ; 2 ♂♂, same locality, 27.10.1881, Feiller leg. ( NHMW9534 View Materials ) ; 1 specimen, same locality, 15.11.1881, Feiller leg. ( NHMW9535 View Materials ) ; several specimens, same locality, 24.11.1881, Feiller leg. ( NHMW9536 View Materials ) ; 3 specimens, same locality, 12.01.1882, Feiller leg. ( NHMW9537 View Materials ) ; several specimens, same locality, 18.01.1882, Feiller leg. ( NHMW9538 View Materials ) ; 1 specimen, same locality, 31.10.1882, Feiller leg. ( NHMW9539 View Materials ) ; 2 specimens, same locality, 16.11.1882, Feiller leg. ( NHMW9540 View Materials ) ; 1 specimen, same locality, 14.07.1883, Feiller leg. ( NHMW9541 View Materials ) ; 1 ♀, same locality, 23.03.1886, Feiller leg. ( NHMW9542 View Materials ) ; 1 ♂, same locality, 24.10.1886, Feiller leg. ( NHMW9543 View Materials ) ; 11 specimen, C. Attems det., same locality, 26.10.1893, ( NHMW9494 View Materials ) ; 11 ♂♂, C. Attems det., same locality, 1898, ( NHMW9495 View Materials ) ; several ♀♀, Praterau , 26.10.1881, Feiller leg. ( NHMW9533 View Materials ) ; 2 specimens, same locality, 24.09.1886, Feiller leg. ( NHMW9544 View Materials ) ; Ottakring : 10 ♂♂, 2 ♀♀, Heuberg SE valley (W Andergasse ), ca 290-310 m, 31.10.1982, J. Gruber leg. ( NHMW2888 View Materials ) ; 8 ♂♂, 1 ♀, Heuberg valley to Predigtstuhl ( Bieglerhütte ), ca 290-310 m, 05.11.1978, J. Gruber leg. ( NHMW9658 View Materials ) ; 1 ♂, same locality, 21.10.1979, J. Gruber leg. ( NHMW9659 View Materials ) ; 1 ♀, same locality, 14.10.1982, J. Gruber leg. ( NHMW9660 View Materials ) ; Penzing : 2 juveniles, Haltertal (S Spitalwiese), ca 270 m, 02.08.1980, J. Gruber leg. ( NHMW9653 View Materials ) ; 1 ♂, same locality, 18.10.1980, J. Gruber leg. ( NHMW9654 View Materials ) ; 1 ♀, Hinter-Hainbach , ca 350 m, 22.10.1988, J. Gruber leg. ( NHMW9655 View Materials ) ; 1 ♀, Kolbeterberg , NE-slope, ca 300 m, 08.10.1977 - 05.11.1977, J. Gruber leg. ( NHMW9656 View Materials ) ; 3 ♂♂, 1 ♀, same locality, 15.11.1977 - 10.12.1977, J. Gruber leg. ( NHMW9657 View Materials ) ; Währing : 1 ♂, Heuberg NE valley (W Andergasse ), ca 400 m, 01.11.1977, J. Gruber leg. ( NHMW9661 View Materials ) ; 2 ♂♂, 5 ♀♀, same locality, 25.12.1980, J. Gruber leg. ( NHMW9662 View Materials ) . GERMANY: Saxony : 1 ♂, Röttis , 03.10.2014, H. Reip leg. (Reip collection); Thuringia : 1 ♂, Laasdorf , 06.11.2007, H. Reip leg. (Reip collec- tion) ; 3 ♂♂, 2 ♀♀, same locality, 26.10.2010, H. Reip leg. (Reip collection) ; 1 ♀, H. Harald det., Windknollen , 17.10.2005, H. Reip leg. (Reip collection) ; 1 ♂, 1 ♀, Zeitzgrund , 17.10.2014, H. Reip leg. (Reip collection). SWIT- ZERLAND : 1 ♂, St. Gallen , 14.09.2013, H. Reip leg. (Reip collection) .

Distribution ( Figs 38 View FIGURE 38 , 39 View FIGURE 39 ). Austria: Ramsach; Buchheimer Tobel; Kristberg ( Meyer & Singer 1997); Rheintal ( Mathis 1951); Dünserberg ( Schied 2012); near Innsbruck ( Thaler et al. 1987); Antelsberg ( Thaler 2003); Kranebitter Klamm, Hechenberg ( Thaler 1982); Rinn; Hochriess ( Kurnik 1988); St. Gilgen; Ischl; Obertraun; Schafbergalpe; Berghang near Admont (Verhoeff 1930); Almkogel; Ardning-Frauenberg; Dürrenschöberl; Admont: Stiftsgarten, Moorwirtschaft, Hall; near Lunz (Attems 1954, present study); Vorau; Spital near Tauchen; Trübenbach; Puchenstuben; Kirchberg an der Pielach; Türkenloch near Kleinzell; Dürre Wand; Hermannshöhle; Krumbach; Prater; Lobau; Hundsheimer Berg ( Attems 1949); Gaisberg; St. Anton am Arlberg; Rosannaschlucht ( Verhoeff 1929a); Gesäuse; Mariazell; Bürgeralpe; Erlaufsee; Wiener Schneeberg; Hainfeld; Araburg; Puchberg; Bregenz ( Verhoeff 1929b); Rothwald; Ötscher-Krummholzstufe; Purgstall; Schauboden ( Ressl 1995); numerous localities near Vienna; Kierling Ufer; Gaming; Kaltenleutgeben; Lilienfeld; Weitenegg; Amstetten; Dunkelsteinerwald; Kemmelbach; Türnitz; Hilm; Triesting; Burgenland; Weingraben; Perschenegg; Mank-Auen; St. Pölten; Hausleiten; Obergrafen— Hubertendorf; Baden; Grimmenstein; Ysperklamm; Hainfeld; Steinwandgraben; Böheim St. Aegyd; Plosdorf; Zauchbach; Traismauer; Hainburger Berge (present study). Czech Republic: Blatná; Rtyně v Podkrkonoší; Hrubý Jeseník; Lednice ( Lang 1971, all four localities are with questionmark); Horní Jiřetín ( Tajovský et al. 2014a); Chvojnov; Na Horních Jemničkách; Špičák ( Tajovský 2017a); Čížov; Konopiště (Kocourek 2018); Jordán; Klobouček; Skelná Huť ( Kocourek & Tajovský 2018). All of the following localities are published within the map in Kocourek et al. (2017) (for details see Schubart 1934; Kocourek 1973, 2013, 2014; Kocourek & Tajovský 2011; Kocourek et al. 2000; Tajovský 2001; Skoumalová 2010; Verhoeff 1927): Kubova Huť; Branov; Týřovice; Týnec nad Sázavou; Hejnice; Nové Město pod Smrkem; Chrášťany (Rakovník); Přílepy (Rakovník); Chyňava; Domoušice; Klánovice; Kojetice; Kuři; Železná Ruda; Horní Hanychov, Liberec; Loučná; Hracholusky (Rakovník); Mladé Buky; Mlýny (Děčín); Stará Oleška (Děčín); Nyrsko; Kunratice; Písnice; Běchovice; Nová Pec; Troskovice; Oldiichov v Ha- jich; Stará Chodovská; Roztoky; Srbsko; Pařízek; Černé Voděrady; Dívčí kámen; Rožnov pod Radhoštěm. France: Vosges, Klintzkopf ( Bigler 1913). Germany: Pappritz (Dresden); Radeberg; Geising (Altenberg); Dohna; Niederwartha (Dresden) ( Verhoeff 1910b); Greiz; Jocketa (Pöhl) ( Verhoeff 1917); Dietersbachalpe; Warmatgundalpe ( Verhoeff 1929b); Erzgebirge; Landkreis Waldshut; Zella-Mehlis; Landkreis Oberallgäu; Plauen ( Schubart 1934, present study); Albbruck; Schlüchttal ( Verhoeff 1936a); Marquartstein ( Verhoeff 1941); Zwickau; Wulm; Schlunzig; Annaberg-Buchholz, Bärenstein ( Büttner 1963); Gutenberger Höhle ( Dobat 1963); Oberlausitz, Kippenstandorte ( Dunger 1966); Osterzgebirge ( Richter 1967); Laichingen ( Dobat 1975); Loch (Königstein); Welluck; Nonnhof (Alfeld); Saass; Raitenbuch (Berching); Hartenstein (Mittelfranken); Bernheck (Plech) ( Dobat 1978); Schönau am Königssee ( Boller 1986); Sächsische Schweiz ( Hauser & Voigtländer 2008); Zeitzgrund (present study); All of the following localities are published within the map in Hauser & Voigtländer (2019) (for details see Reip et al. 2012): Ottendorf;Abbachhof (Wenzenbach); Eurasburg (Oberbayern); Eibenstock, Weitersglashütte; Dresden, Geising (Altenberg); Müglitztal; Oberwiesenthal; Tannenbergstahl; Frauenau; Zwieslerwaldhaus; Lam; Drachselsried; Frauenstein; Schlüchtthal; Harz; Allgäu, Staufenberg; Jena; Schönbrunn; Bad Lobenstein; Bruck an der Amper; Mühltal; Planegg; Kreuzeck; Reschbachklause; Latschenkopf; Riedl; Nebelhorn; Bodensee; Bankholzen; Ruine Rosenegg; Ratzenried; Grosser Arber; Rachelsee. Hungary: Násznép Cave, Naszály Mt. ( Loksa 1959); Pilis Mt. (Loksa 1991). Luxembourg: three caves ( Weber 2013). Poland: the only certain record is from Góra Połom, Wojcieszów (Bielak- Oleksy & Jaskiewicz 1977); Tatry Mountains ( Stojałowska & Staręga 1974). Slovakia: Little Carpathians, near Bratislava ( Gulička 1956); Žilinský lesopark ( Stasiov & Hazuchova 2010). Switzerland: Rochers de Naye ( Verhoeff 1894); Schweizer Blauen; Guldental ( Bigler 1913); Val Cluoza; Selva Zernez ( Bigler 1929); Tinière ( Faës 1902); Breitlauenen, Berner Oberland ( Rothenbühler 1899); Bremgartenwald, Bern ( Rothenbühler 1900); St. Gallen (present study); in addition to those, there are several more localities mapped in Swiss Plateau; Grisons, Rhine basin; Engadin; northern slope of the Alps (see Pedroli-Christen 1993: XXXVI).

Type locality. Austria-Hungary, unknown locality.

Comments. Since H. flavescens males with 28 body segments are known (see Spelda 2005) and since the coxal process of the posterior gonopods show some variability in Haasea species, we synonymize here Orobainosoma flavescens vornatscheri Verhoeff, 1935 with Haasea flavescens . Strasser (1966: 183) already stated a probable synonymy between vornatscheri and pinivaga (now flavescens ), but he did not officially synonymize them.

The examination of the Latzel’s type series of Craspedosoma flavescens housed in NHMW (see Wirkner et al. 2002: 13) revealed that the only two males assigned to this species are actually H. germanica and H. fonticulorum , respectively. At present, the male described by Latzel remains known only from the original description, unless the illustrated gonopod actually corresponds to a lose anterior gonopod not included in the type series of H. flavescens (NHMW9549) and part of the numerous fragments and juvenile specimens of Latzel’s collection of the species subsequently received by NHMW in 1919 and revised by J. Gruber in 2007. Although this material is probably part of Latzel’s type series acquisitioned in NHMW, we preferred not to designate a lectotype for the species based on this fragment.

In his original description of C. flavescens, Latzel (1884) stated he examined around 80 specimens, mainly juveniles from “Salzburg, Oberösterreich, Niederösterreich, Westungarn und Kärnten” as well as one specimen from “Tirol, Krain, Croatien, Galizien und Oberungarn” and France. Doubt in such a distribution of H. flavescens was already expressed by Verhoeff (1907: 326), who stated that these were probably several species.

Attems (1895, 1929, 1949, 1954) contributed to this confusion by assigning several known and undescribed taxa to Orobainosoma flavescens (see Fig. 10 View FIGURE 10 and comments for H. cyanopida , H. filicis , H. fonticulorum , H. ger- manica, H. hungarica , H. inflata , H. plasana and H. gruberi sp. nov.), which in its turn generated mistakes in understanding species occurrences and distributions. In fact, Strasser & Minelli (1984) recognized that Attems’ (1949) record of H. flavescens from “ Lombardy ” in Italy as dubious. Probably this was H. fonticulorum . The record of O. flavescens from Bachern, Marburn (Maribor), based on 19 juveniles ( Attems 1949) is here assigned to Haasea inflata . Based on these facts, we delete H. flavescens from the list of species of Croatia, Bosnia and Herzegovina, Slovenia and Italy. Two records of O. flavescens from Reistal (= Reisthal Rax) and Schöckl in Austria by Attems (1949), are not included in the distribution map here since these were based on juveniles and females, and four more species occur in the region.

Additionally, records of H. flavescens from the Czech Republic based on Lang’s (1971) unpublished monograph should be taken with reserve, since these could also be based on misidentified specimens (Dolejš, pers. comm.). The same is true for the Luxembourg records by Weber (2013), which could refer instead to the genus Xylophageuma . These localities are marked with question marks although we do not exclude that they might be good identifications. The record from Vosges Mt. in France by Bigler (1913) is considered correct since he described the second species of the genus Xylophageuma , leaving little chance that he could have mixed these species. Bröleman’s (1935) record of H. flavescens from French Jura was probably referred to Bigler’s (1913) record from Swiss Jura, so we exclude it here though the occurrence of flavescens in Jura Mountain in France is also highly plausable. The record from Tatry Mountains should be also taken with reserve as it could refer to Hylebainosoma .