Pterostichus

Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin & Zhou, Hongzhang, 2013, A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions, Zootaxa 3664 (2), pp. 101-135 : 103-104

Key to subgenera of the genus Pterostichus from China (part)

1. Mesofemora with two or three setae on posterior margin; metaepisterna short, length subequal to the width of anterior margin; metacoxa with only two setae; hind trochanter without setae; penultimate labial palpomere without seta near apex; pronotum

with hind angles distinct, posterior seta placed near hind angle.................................................. 2 - Without the combination of above characters.......................................... Pterostichus other subgenera

2. Hind trochanter without seta............................................................................. 3

- Hind trochanter with one seta...................................................... Pterostichus other subgenera

3. Third interval of elytron with three or more setigerous pores, 5th interval without pores; ridge between outer basal fovea of pronotum and lateral margin strongly carinate; right paramere thick.................. Pterostichus ( Feroperis Lafer 1979 )

- Third interval of elytron usually with two or less setigerous pores; if 3rd interval has more than three pores, then 5th interval has some pores; ridge between outer pronotal basal and lateral margin strongly or at most moderately convex; right paramere lamina............................................................................................. 4

4. Apical lamella of male genitalia laminar, rather wide and long, usually slightly twisted; species from the Himalayan Region....................................................................... Pterostichus ( Pseudethira Sciaky, 1996 )

- Male genitalia not like so; species not from the Himalayan Region............................................... 5

5. Umbilical series on 9th interval continuous; apical lamella of male genitalia long and strongly twisted................................................................................... Pterostichus prattii Bates, 1890 species group

- Umbilical series on 9th interval discontinuous; apical lamella of male genitalia at most slightly twisted.................. 6

6. Apical lamella of male genitalia wide, more or less bent dorsally; pronotal disc impunctate; females with strong isodiametric microsculpture on elytra; setae on terminal sternum distant from apex................. Pterostichus (Wraseiellus n. subg.)

- Apical lamella of male genitalia narrow and short, not bent dorsally; pronotum densely and finely punctate; setae on terminal sternum close to apex. (only applicable for Nialoe species from China)............... Pterostichus ( Nialoe Tanaka, 1958 )