Systaria panay, Jäger, 2018

Jäger, Peter, 2018, On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests, Zootaxa 4504 (4), pp. 524-544 : 537

Systaria panay spec. nov.

Figs 31–32 , 45–46

Type material: PHILIPPINES: Visayas : Holotype: female, NW Panay Island , PESCP Research Station, Sibaliv, 11°49'11''N, 121°58'0''E, 450 m elevation, Cichosz & Einhäupl leg. 9.2008 ( SMF).

Etymology. The species name is derived from the type locality; term in apposition.

Diagnosis. Small spiders with body length of females: 6.6. Females are very similar to those of S. deelemanae Dankittipakul & Singtripop 2011 in having a similar conformation of the atrial rims, the position of copulatory openings and the course and shape of the internal duct system especially the loop of copulatory ducts close to the copulatory openings ( Figs 31–33 ), but may be distinguished by the distinctly wider distance between the copulatory openings (closer in S. deelemanae ), the epigynal field wider than long (distinctly longer than wide in S. deelemanae ) and the shorter and stouter bursae (strongly elongated in S. deelemanae ).

Description. Female (holotype): prosoma length 2.9, prosoma width 1.9, anterior width of prosoma 1.2, opisthosoma length 3.7, opisthosoma width 2.0. Eyes: AME 0.18, ALE 0.13, PME 0.15, PLE 0.15, AME–AME 0.08, AME–ALE 0.03, PME–PME 0.19, PME–PLE 0.15, AME–PME 0.10, ALE–PLE 0.02. Cheliceral furrow with 3 promarginal and 2 retromarginal teeth. Spination: Palp: femur p001, d011, patella [only with thin setae], tibia p21, d11, metatarsus p21, d10, r10, v012; legs: femur I p0011, d111, II p001, d111, r010[small], III p111, d11(2)1, r111, IV p101(110), d111, r001; patella I–IV 000; tibia I v21 (0)0, II v100, III p11, d010, r11, v212, IV p11, d010, r 11, v22 (3)2; metatarsus I–II v100, III p112, r112, v22 (1)1, IV p122(212), r21(2)2, v2121. Leg formula: 4123. Measurements of palp and legs: Palp 3.3 (1.1, 0.5, 0.7, -, 1.0), I 8.1 (2.2, 1.0, 2.1, 1.7, 1.1), II 7.9 (2.1, 1.0, 2.0, 1.7, 1.1), III 7.3 (2.0, 0.9, 1.6, 1.8, 1.0), IV 10.0 (2.6, 1.0, 2.5, 2.7, 1.2). Copulatory organ as in diagnosis ( Figs 31–32 ). Epigynal field pointed anteriorly, with 1 pair of slit sensilla, without anterior bands. Atrium roughly pentagonal with anterior and posterior margins indistinct and two indistinct light windows in the centre. Lateral atrial rims converging posteriorly, with distinct bend anteriorly. No sclerotised spheres recognisable. Bursae slightly longer than wide, separated by their average width. Spermathecae separated by about 3 of their diameters, fertilisation ducts arising medially, anteriad. Colouration. Pale yellowish-brown without distinct pattern, labium, gnathocoxae, margins of sternum, eye region and fovea darker. Femora and patellae with indistinct dark grey patches. Opisthosoma greyish, dorsally with light longitudinal stripe in anterior half, ventrally with dark spinning glands visible through cuticle in front of spinnerets.

Male: unknown.

Distribution. Known only from the type locality ( Figs 45–46 ).

Natural history. Spiders have been recorded from an elevation of 450 m, obviously found in the forest.

Figures

FIGURE 45. Records of Systaria spp. in SE Asia described in this paper. Triangles—species recorded from caves, circlesspecies recorded from forest.

FIGURE 46. All known records of the genus Systaria (World Spider Catalog 2018) including undescribed species mentioned in Deeleman-Reinhold (2001) according to habitats. Triangles—species recorded from caves, circles—species recorded from forest. See discussion.

FIGURES 27–32. Systaria spp., copulatory organs. 27–28 Systaria bregibec spec. nov., holotype male from Cambodia, Rabbit Island. 29–30 Systaria bifidops spec. nov., holotype male from Malaysia, Bukit Fraser. 31–32 Systaria panay spec. nov., holotype female from Philippines, Panay Island. 27–30 Left palp (27, 29 ventral, 28, 30 retrolateral). 31 Epigyne, ventral. 32 Vulva, dorsal. Abbreviations: BP—bifid process of RTA, CO—copulatory opening, LO—loop of copulatory ducts, TPproximal projection of tegulum, VH—ventral hump of palpal tibia.

FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening.