Systaria lannops, Jäger, 2018

Jäger, Peter, 2018, On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests, Zootaxa 4504 (4), pp. 524-544 : 525

Genus Systaria Simon 1897

Note. The terminology of the female internal duct system was used ambiguously in previous literature. Copulatory duct (part, in which embolus is inserted; insemination duct sensu Dankittipakul & Singtripop 2011) connects the copulatory opening with the part of the duct system where the bursae branches off. The part between that point and the spermathecae I call connecting tube according to the terminology used by Jin et al. (2016) for the genus Otacilia Thorell 1897 ( Phrurolithidae ). Both ducts might be short and hardly discernible. As in Jäger (2005: 88), slit sense organs close to the epigyne are illustrated as descriptive characters. Same is true for sclerotised spheres in the epigyne (see Jäger 2012). These were interpreted as remnants of fused epigynal furrows. Parts of these furrows can be seen in Systaria lannops spec. nov. and S. procera spec. nov. In S. longinqua spec. nov. and S. princesa spec. nov. such furrows are absent and generally more spheres are present then (see also discussion). The sternum of all Systaria species treated in this paper exhibit 4 pairs of indistinct humps, the posterior one almost fused. Between the humps 3 pairs of slit sensilla on each side are present, each consisting of 1 large and 1 smaller slit sensillum ( Fig. 35 ). The distal half of metatarsi and tarsi exhibit sparse scopula ventrally, spinnerets have long setae, posterior lateral spinnerets exhibit the distal segment distinctly longer than in remaining spinnerets ( Fig. 15 ). Claw tufts are dense.

Figures

FIGURES 33–39. Systaria princesa spec. nov. from Philippines, Palawan (33–34, 37 holotype male from Tagusan cave, 35– 36 male paratype from Puerto Princesa Underground River cave, 38–39 female paratype from Tagusan Cave). 33–36 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening.

FIGURES 11–15. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening.