Thinodromus makarovi, 2020

Gildenkov, M. Yu., 2020, Three new species of the genus Thinodromus Kraatz, 1857 from Indonesia and the Philippines (Coleoptera: Staphylinidae: Oxytelinae), Russian Entomological Journal 29 (2), pp. 148-152 : 149-151

Thinodromus (s.str.) makarovi Gildenkov , sp.n.

Figs 2 , 7–9 .

MATERIAL. Holotype, ♂, Indonesia, West Java: with labels “ INDONESIA: W Java Gn. Salak, 8km S Bogor Sungai Ciapus, ca 800m lg. Schuh 17.8. 1994 ” ( NHMW) . Paratypes: 3♂♂, 1♀ “ INDONESIA: W Java Gn. Salak, 8km S Bogor Sungai Ciapus , ca 800m lg. Schuh 17.8. 1994 ” ( NHMW; 1♂ — cMG) ; 2♂♂, 1♀, Indonesia, West Sumatra: with labels “INDONESIEN 1991 (28) W-Sumatra, W Danau di Atas 70 km SE Padang, leg. Jäch 24.2” ( NHMW) ; 4♂♂, 5♀♀, 5 ex., Indonesia, North Sumatra: with labels “ SUMATRA ACEH Ketambe 7.–17.X.> < INDONESIA 1991, leg. Barries ” ( NHMW; 1♂, 1♀, 1 ex. — cMG) ; 1♀, 2 ex. “ SUMATRA, Aceh Kutacane. Ketambe 7.- 19.10.1991, 400m, leg. Barries ” ( NHMW) .

DESCRIPTION (holotype). Length 3.3 mm. Colouration brown, legs light-brown. Integument slightly shining, body with short, light-coloured seta. Some specimens are blackbrown, legs and antennas are brown.

Head transverse, with a wide base, ratio of its length (from posterior margin of head to anterior margin of clypeus) to maximum width is about 28:43. Neck constriction prominent. Eyes large, convex, occupying almost entire lateral side of head; temples almost indistinct, the head has the largest width in the eye area ( Fig. 2 ). Head surface with rather clearly, highly finely and dense punctation. The diameter of the punctures is more than 5 times smaller than the diameter of the eye facet. Distances between punctures significantly smaller than their diameter, interspaces smooth, slightly shining ( Fig. 2 ). Antennae rather long, antennal segments 1–8 elongated; segments 9–10 are about as long as wide; segment 11 elongated, conical ( Fig. 2 ).

Pronotum heart-shaped, reaches its greatest width after about 2/3 of the length measured from the base, then narrowed ( Fig. 2 ). Ratio of pronotum length to its maximum width about 37:46. Surface of pronotum with rather clearly, finely and dense punctation. The diameter of the punctures is approximately 5 times smaller than the diameter of the eye facet. The distance between punctures is significantly smaller than their diameter, interspaces smooth, slightly shining ( Fig. 2 ). Base of pronotal disc with distinct horseshoe-shaped depression ( Fig. 2 ); central part of disc with 2 symmetrical distinct weakly developed oval depressions.

Ratio of length of elytra to their combined width about 65:76. Elytra with rather clearly, finely and dense punctation. The diameter of the punctures is about 4 times smaller than the diameter of the eye facet. Distance between punctures are a little smaller than their diameter, interspaces smooth, shining ( Fig. 2 ).

Abdomen with fine and dense punctation, shining.

Aedeagus of characteristic structure ( Figs 7–8 ).

Female. Sexual dimorphism is absent, female morphologically similar to male. Spermatheca of characteristic structure ( Fig. 9 ).

COMPARATIVE REMARKS. The species is distinguished by large sizes, well- developed eyes, and distinct, fine, and dense punctation of the body surface. Reliably different in the structure of the aedeagus ( Figs 7–8 ).

DISTRIBUTION. Indonesia: Java, Sumatra ( Fig. 14 ) .

ETYMOLOGY.The species is named after Kirill Vladimirovich Makarov, a well-known expert in the biology and taxonomy of Coleoptera .

13

Figures

Figs 1–3. Thinodromus spp., holotypes, males, dorsal view: 1 — Thinodromus (s. str.) coronatus, sp.n.; 2 — Thinodromus (s. str.) makarovi, sp.n.; 3 — Thinodromus (s. str.) shavrini, sp.n. Рис. 1–3. Thinodromus spp., голотипы, самцы, сверху: 1 — Thinodromus (s. str.) coronatus, sp.n.; 2 — Thinodromus (s. str.) makarovi, sp.n.; 3 — Thinodromus (s. str.) shavrini, sp.n.

Figs 4–12. Genitalia of Thinodromus: 4–6 — T. coronatus, sp.n.; 7–9 — T. makarovi, sp.n.; 10–12 — T. shavrini, sp.n.; 4, 7, 10 — aedeagus, ventral view (holotypes); 5, 8, 11 — aedeagus, lateral view (holotypes); 6, 9, 12 — spermatheca (paratypes). Scale bars: 0.25 mm. Рис. 4–12. Гениталии Thinodromus: 4–6 — T. coronatus, sp.n.; 7–9 — T. makarovi, sp.n.; 10–12 — T. shavrini, sp.n.; 4, 7, 10 — Эдеагус, сниЗу (голотипы); 5, 8, 11— Эдеагус, сбоку (голотипы); 6, 9, 12 — сперматека (паратипы). МасШтаб: 0,25 мм.

Figs 13–14. Distribution of Thinodromus: 13 — T. coronatus, sp.n. (black circles) and T. shavrini, sp.n. (black squares) in Philippines; 14 — T. makarovi, sp.n. in Indonesia. Рис. 13–14. Распространение Thinodromus: 13 — T. coronatus, sp.n. (черные круги) и T. shavrini, sp.n. (черные квадраты) на Филиппинах; 14 — T. makarovi, sp.n. в ИндонеЗии.