Trichogenia ulmeri, Webb & Braasch & Mccaferty, 2006

Webb, J. M., Braasch, D. & Mccaferty, W. P., 2006, Reevaluation of the Genera Compsoneuria Eaton and Trichogenia Braasch & Soldán (Ephemeroptera: Heptageniidae), Zootaxa 1335, pp. 55-68 : 60-61

Trichogenia Braasch & Soldán

As indicated above, the larva Ulmer (1939:675) described as C. spectabilis is actually an unnamed species of Trichogenia [= Trichogenia ulmeri N.SP.]. Wang and McCafferty (2004) considered Ulmer's C ompsoneuria to belong to a different genus than Trichogenia based on differences in mouthpart morphology. However, examination of a new species described below and T. ulmeri shows that these two species are fundamentally the same as T. maxillaris Braasch and Soldán, 1988 and differ in only a few characters. Therefore, we place them all in Trichogenia .

Adult males have not been associated with any larvae of Trichogenia . However, it is highly probable that Heptagenia nasuta Ulmer, 1939 , known only from adults from Indonesia, is Trichogenia ( Braasch and Soldán 1988) based on the following: i) the only confirmed larvae of the subfamily Heptageniinae known from southeast Asia are Trichogenia ; ii) H. nasuta is the only adult of the subfamily Heptageniinae known from southeast Asia; iii) wings dissected from wingpads of mature Trichogenia larvae have a slight violet tinge, similar to H. nasuta ( Braasch and Soldán 1988) . For these reasons we provisionally transfer H. nasuta to Trichogenia [= Trichogenia nasuta (Ulmer) N.COMB.].

Diagnosis: Larvae of Trichogenia can be differentiated from those of all other genera of Heptageniidae by the following combination of characters: i) ventral side of maxillae with setae in a row; ii) tergum with branched robust setae and many long, fine setae; iii) lamellae of gills 1 reduced; iv) lamellae of at least gills 2-4 long, narrow, and pointed apically; v) apex of femora without dorsal projection.

At this time we do not have material available to provide characters for separating adult males of Trichogenia from other Heptageniinae genera. The long pair of ventral spines on the penes shown in Figures 156 and 157 by Ulmer (1939:569) in combination with the violet coloration in the wings may be diagnostic.

Species included: Trichogenia maxillaris , T. nasuta N.COMB., T. hubleyi N.SP., T. ulmeri N.SP.