Ancyropilus simplex, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 19-21

Ancyropilus simplex sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:FA9C15ED-CF77-4BF3-A033-E401928ACE3E

( Figs 33 , 72 , 108 , 146 ; Map 1)

HOLOTYPE (sex not determined): Western Australia: 22.09S 118.07E GPS, 25km NNW of Wittenoom Gorge , WA, 4 Oct. 1995, D.C.F.Rentz & L.Lowe, Stop 92 ( ANIC) . PARATYPES (42): Western Australia: same data as holotype (1 ♀, 1, ANIC); 25 km NW Bandya Stn. , W. Aust., 2.xi.1989, K.T. Richards // Agriculture (Dept.) Western Australia 49857 (1 ♁, WADA); Carnarvon WA., 25-II-77, M.S. Moulds // Opilo congruus Newm. A. Walford Huggins det. 1983 (1 ♀, QM); 107 miles SSE of Carnarvon , WA, 21 April 1968, I.F.B. Common & M.S. Upton (1 ♀, 4, ANIC); Australia, WA Menzies , 10 km North , 14 January 1989, M.S. Moulds // K 304506 (1, AM); WA, Pilbara region. Juna Downs Station, Great Northern Highway , approx. 8km S of Karijini Dr toff., -22:41:36, 118:42:19, 3- Jan-2006 to 8-Jan-2006, LTM sites, A. Donnelly & CVA Volunteer PILB038/11M (Malaise trap) // 1750 // K 246911 (1, AM); WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28; 16- Feb-2005, 20-Feb-2005, LTM sites, M. Bulbert & S. Ginn, PILB044/04M // 1135 // K 246910 (1, AM); WA, Pilbara region. Hamersley station, Horseshoe Bore , 300m N of windmill, -22:29:10, 117:41:28, 3-Jan-2006 to 8-Jan-2006, LTM sites, A, Donnelly & CVA volunteer, PILB044/11M (Malaise trap) // +2 // 1749 // K 246912 (1, WAM E113528 , ex AM); WA, Pilbara region. Juna Downs Station, Juna Downs Rd to Packsaddle Bore approx. 5km E of homestead., -22:52:31, 118:31:49, 19-Nov-2004 to 24-Nov-2004, LTM sites, CVA volunteers, PILB039/03M (Malaise trap) // 1140 // K 246905 (1 ♁, AM); WA, Pilbara region . Juna Downs Station , Juna Downs Rd to Packsaddle Bore approx. 20km E of homestead, -22:51:30, 118:40:14, 3-Jan-2006 to 8-Jan-2006, LTM sites, A. Donnelly & CVA volunteer, PILB036/11M (Malaise trap) // 1753 // K 246898 (1, WAM E113529 , ex AM); WA, Pilbara region . NanburraWittenoom Rd , approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 18-Nov-2004 to 23-Nov-2004, LTM sites, CVA volunteers, PILB041/03M (Malaise trap) // 1137 // K 246908 (1, AM) ; same data as previous except: 1138 // K 246907 (1 ♁, WAM E113530 , ex AM) ; same data as previous except: 1139 // K 246906 (1 ♁, WAM E113531 , ex AM); WA, Pilbara region. Nanburra-Wittenoom Rd , approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 28-Oct-2005 to 2-Nov-2005, LTM sites, CVA volunteers, PILB041/10M (Malaise trap) // 2030 // K 246904 (1, AM); same data as previous except: 2031 // K 246913 (1, WAM E113532 , ex AM); WA, Pilbara region. Nanburra-Wittenoom Rd , approx 13km NE of Railway Rd. x-ing, nr fenceline, -22:26:8, 117:49:56, 13-April-2005 to 18-April-2005, M. Bulbert & G. Wood, CVA volunteers, PILB041/05M (Malaise trap) // 1136 // K 246909 (1, AM); 28.22S 122.37E Deeba Rock Hole 34km NEbyE of Laverton WA 12.xi.1977 T.A. Weir (1, ANIC); Marloo Stn. Wurarga, W.A. 1931-1941 A. Goerling (1 ♁, 3 ♀, 6, ANIC); 7.5 km SE of Banjiwarn HS (27°42′S 121°37′E) W. Aust. 22-28 Feb. 1980. T.F. Houston et al. 316-10 // at light at night // Western Australian Museum Dept. of Biological Survey Site. B W Camp (1, WAM E88383 ); 37 km SW Youanmi. Western Australia. 28.45 S 119.31 E. 13-14 March 1982. T.F. Houston & B. Hanich. 437-8 // at light at night (1, WAM E88384 ); Kathleen Valley 27°24′S 120°39′E Western Australia 1963 T. Moriarty (1, WAM E88386 ); WA Charles Darwin Reserve, 2.5 km N White Wells HS 29°33′39″S 116°57′52″E 4-5 March 2008 T.F. Houston 1254-2b // ex gas lantern / pitfall trap 10:00pm to dawn, mixed shrubland with emergent eucalypts and native pine (1, WAM E88387 ); Ca. 1 km WSW of Koora Retreat, Koorarawalyee , Western Australia. 31°15′54″S 120°00′50″E 26-27 December 2011 T.F. Houston 1390-1 // Ex fluoro-light/ pitfall trap operated 7:45 to 4 am (1, WAM E88388 ); Mogumber , 16.xii.55 , Collector R.P. McMillan (1, WAM E88385 ); Wooramel River, West. Australia 25°47′S 115°58′E 14 April 1979, M. Peterson // at light at night (1, WAM E88380 ); Warnie River , 49 km E of Paynes Find , Western Australia, 12-13 March 1982, T.F. Houston & B. Hanich, 436-1 // at light at night (2, WAM E88374 , E88375 ) ; Wannara Stn [ Wannarra Station ] WA, 13 Feb 2010, Light, SR Patterson (1, WAM E88401 ) .

Diagnosis. Pronotum robust, disc very densely punctate except narrow longitudinal stripe from central impression to pronotal furrow, and small obscure area either side of central impression, lacking punctation; elytron blackishbrown with pale orange fasciate maculation, fascia rounded before suture, distinct humeral and apical maculae absent (common) or small apical macula present (uncommon), punctation lacking nodules, 8 th stria beginning anterior to fascia, between 3 rd and 8 th punctation of 7 th stria, all striae almost complete to apex; femora entirely dark brown, only profemora slightly swollen; tarsi with three ventral tarsal pads.

Ancyropilus simplex sp. nov. is most similar to A. noonbahensis sp. nov. though differs from it by the tegmen with base of parameroid lobes weakly chitonised externally and dorsal sinus V-shaped, the femora entirely black, and the apical elytral maculation absent (common) or only faintly represented (uncommon).

Description. Habitus: Fig. 146 . Total length: 7.3–13.9 mm (holotype, 8 mm). Head: Cranium black, clypeus and supra-antennal elevations reddish-black, anteclypeus semi-transparent orange, labrum, palpi and antennae orange to reddish-brown; vertex and frons with dense network of irregularly-shaped punctation, above clypeus with weak transverse rugulosity; clypeus variable, rugulose to punctate, anterior margin straight to weakly angulateconcave; eyes separated by 0.65–0.96 eye widths (holotype, 0.75); genae wrinkled; terminal maxillary palpomeres with exterior margins about 1.8 times the length of inside edges; terminal labial palpomeres with exterior margins about 2.5 times the length of inside edges; antennae almost reaching base of pronotum; setae pale, long erect setae on, and bordering, eyes and along clypeus, shorter medially-directed on frons, posteriorly-directed setae above eyes. Prothorax: Black to blackish-brown; pronotum 1.13–1.24 times longer than wide (holotype, 1.19), middle about as wide as pronotal arch, sides rounded, lateral impression obscure, subapical depression v-shaped, central discal impression shallow, indistinct; disc very densely punctate except a conspicuous slender impunctate longitudinal line from central impression to pronotal collar and small obscure area before pronotal arch either side of central impression, pronotal arch and collar less densely punctate; setae pale, dense fine multi-directional setae forming swirld around impunctate areas, thicker longer erect setae most dense laterally. Pterothorax: Ventrites black or brownish, metaventrite with seta-associated punctation, setae short pale posteriorly-directed; elytra blackish-brown with a single pale-orange maculation (each elytron with a broad fasciate band rounded at the suture), apices lacking maculation (common) or with a small pale macula (uncommon); length to width ratio 2.73–2.91:1 (holotype, 2.75); punctation circular, large anterior to fascia, smaller posterior of fascia, without internal nodules, all striae terminating near apex, 8 th stria beginning anterior to fascia, between 3 rd and 8 th punctation of 7 th stria (beginning closer to base in larger-sized specimens); interstriae about as wide as corresponding punctures (may appear narrower), with rows of fine semi-erect setae (> 1/puncture) and longer erect setae every few punctures, more than one seta often across interstrial width, epipleurae extending into apical curve; hindwing with CuA 3+4 and CuA 1 cross-veins complete, MP 3+4 substantialy extended basad of CuA 1 crossvein. Legs: Femora entirely dark brown, tibiae and tarsi lighter brown, longitudinal tibial carina often darker than rest of tibia, ventral tarsal pads pale brown; profemora slightly swollen, other femora slenderer. Abdomen: Ventrites orange to orange-brown. Male genitalia: Tegmenal plate ( Fig. 33 ) broadest at base, narrowing before parameroid lobes, base of parameroid lobes weakly chitonised externally, tapered and digitiform apically, dorsal sinus about one-quarter tegmen length, wide open externally, internal limit V-shaped, opening at mid-point with short membranous (setose?) lobes, ventral sinus of similar length, apodeme about one-quarter tegmen length; median lobe as in Fig. 72 ; pygidium as in Fig. 108 .

Etymology. The specific epithet, simplex (Latin, meaning single), refers to the single large orange maculation just posterior of the middle on each elytron. This is the only species of the genus in which most indivuals lack maculation on the elytral apices (a more-or-less distinct mark was observed on a single specimen).

Variation. Each elytron of one specimen collected at Juna Downs Station in the Pilbara region has a more-orless distinct, small, pale, apical macula.

Biology. Specimens have been collected from October to April, some using a Malaise trap.

Distribution (Map 1). Ancyropilus simplex sp. nov. is known only from central Western Australia.

Figures

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 138–149. Habitus: (138) Ancyropilus brigalowae, paratype; (139) Ancyropilus emmotti, holotype; (140) Ancyropilus exossuarius, paratype; (141) Ancyropilus hornensis, paratype; (142) Ancyropilus labris, holotype; (143) Ancyropilus monteithi, paratype; (144) Ancyropilus noonbahensis, holotype; (145) Ancyropilus packsaddlensis, holotype; (146) Ancyropilus simplex, paratype; (147) Ancyropilus tricolor, paratype; (148) Culcipternotus mareebensis, holotype; (149) Infectostriatus absentis, holotype (scale bars: 1.0 mm).